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Slender beardtongue (aka, lilac penstemon) is another beardtongue/penstemon that is uncommon in Jackson Park. The stems are unbranched, green or purple, and may or may not bear hairs. There are both basal and stem leaves. The basal and lower stem leaves are 1-3" long and 1/2" wide, variable in shape and leaf margins; the stem leaves are smaller and minutely toothed. Individual flowers are pinkish, purple, or pale violet, about 3/4" long; they are comprised of a green calyx with five triangular lobes and a tubular corolla that is abruptly inflated in the more distal two thirds of the flower. The lower lip of the corolla has three lobes that extend down and out from the aperture the upper lip has two smaller, erect lobes. The interior of the corolla is white with several dark purple lines that act as nectar guides. The filament of a sterile stamen (a "staminode") bearing sparse yellow hairs runs along the midline of the lower half of the corolla, while the filaments of the other four stamens wrap around the inside of the corolla like ribs in a ribcage, with the black anthers positioned on either side of the midline of the upper corolla. A single style runs along the upper midline between the black anthers. The fruit is a more or less bullet-shaped capsule nestled in the persistant calyx. The calyx, flowers, and flower stalks are all covered with glandular hairs; the stems and leaves are not.

Penstemon

Penstemon gracilis

slender beardtongue [Blooms: Jun-Jul]

Hairy beardtongue is 1-2.5' tall and blatantly hairy — backlit plants seen from a distance show a white nimbus around the stems. The flowers are slender, lavender colored, with white lips; the aperture of the flower is nearly occluded by the arched lower lip. Like the other beardtongues/penstemons the tubular flowers have an upper and lower lip have a total of five lobes (two above, three below). The calyx, flower stems, and the flowers themselves are densely covered by glandular hairs; the stems are densely covered with fine white hairs. The stem leaves are pointed, triangular with toothed margins, and occur in opposite pairs; the leaf pairs are oriented at 90° to the pairs above and below them on the stem.

Penstemon

Penstemon hirsutus

hairy beardtongue [Blooms: May-Jul]

Pale beardtongue stems terminate in a 4-10" long panicle of flowers that is longer than broad. Flowers are 3/4" long consisting of a short, green calyx with five teeth, a tubular, white corolla similar to that of other penstemons/beardtongues, four fertile stamens with brown anthers, a bright yellow "beard" on the lower lip (as in hairy beardtongue), and a single style. The upper lip of the corolla has the usual two erect lobes, but the lower lip has three rounded lobes that project forward well beyond the margin of the upper lip. The outer surfaces of the flowers and flower stalks are covered with hairs. The throat of the corolla's lower surface has three thin purple veins, a narrow patch of long, bright yellow hairs, and a pair of low ridges/grooves, one on either side of the midline. The fruit is an egg-shaped capsule 1/4" long nestled in the persistent calyx. Both pale beardtongue and foxglove beardtongue (Penstemon digitalis) have white flowers and might be confused. However, the flowers of pale beardtongue are smaller, the flowers have two distinct ridges on the interior of the lower half of the corolla, and the three lobes on the lower lip are parallel to the floor of the corolla and extend well beyond the upper lip. None of these traits are present in foxglove beardtongue. In Jackson Park, pale beardtongue is less common than hairy beardtongue and much less common that foxglove beardtongue.

Penstemon

Penstemon pallidus

pale beardtongue [Blooms: May-Jun]

Heal-all (a.k.a., self-heal) is a native species superficially similar to ground ivy (Glechoma hederacea) or early-season henbit (Lamium amplexicaule). Heal-all has the characteristics of other low-growing plants in the mint family — an upright, four-angled stem, with lines of hairs along the ridges, 1' tall or less, with a short (1-2") spike of flowers at the apex — and is thus easy to overlook. But the tiny (1/2" long), tubular, two lipped flowers are distinctive (and charming). The light purple or blue upper lip is bent at right angles to form a hood over the aperture with the four stamens and single style with a bifurcated stigma at its tip curving along inside of the hood. The lower lip is divided into three lobes, the middle one white and fringed; the two smaller lateral lobes are the same color as the hood. The hood of the upper lip also sports a "mohawk" haircut of relatively long hairs along the crest of the hood. The flower calyx is light green or reddish, hairy along the edges. After fertilization, four tiny, brown, finely-ridged nutlets develop within the persistant calyx; the seeds are reported to be dispersed by a mechanism that uses the energy in raindrops impacting the calyx to fling the seeds away from the mother plant. The leaves are opposite, up to 2" long and half as wide with a short petiole; they are egg- or lance-shaped with a central vein that is hairy on the underside. Heal-all prefers moist habitats with a history of disturbance.

Prunella

Prunella vulgaris

heal-all [Blooms: Jun-Aug]

American germander flower spikes develop on the tips of the stems, with flowers opening from the bottom of the spike towards the (elongating) tip. Individual flowers are small (about 1/3-3/4"), white or lavender, with a bell-shaped, hairy calyx; an upper lip reduced to two lance-like, vertical lateral lobes that insure pollinators remain beneath the single long, arching single and 3-4 stamens with purple anthers; and a greatly expanded lower lip (a landing pad for pollinators) covered with purple spots. The fruit consists of four nutlets that are nearly fused together, nestled in the remnants of the calyx (which clearly shows the asymmetrical teeth — bottom two smaller than upper three). The nutlets are greenish white when immature, becoming redder and coarsely pitted as they mature. American germander is distinguished from other members of the mint family (Lamiaceae) by the very small upper lip and enlarged lower lip. Related species of Teucrium contain a toxic compound that causes liver damage in animals; for safety's sake, I would avoid herbal remedies that include American germander.

Teucrium

Teucrium canadense

American germander [Blooms: Jun-Aug]

Purpletop vervain flowers occur in dense, flat-topped clusters 1-3" across. Individual flowers are about 1/4" (5-6 mm) long and 8-9 mm long. The flowers contain a short, tubular, hairy calyx with five teeth; a pinkish purple, tubular corolla with five spreading lobes that have notched tips; four stamens which do not extend beyond the throat of the flower; and a pistil with a single style. The fruit is four nutlets, cupped in the remnants of the calyx as they mature.

Verbena

Verbena bonariensis

purpletop vervain [Blooms: Sept]

Prostrate vervain stems produce dense spikes of flowers at their tips. Beneath each flower is a 3/4" long, narrow bract with long, whitish hairs; the bracts make up the erect or sprawling flower spike (6" long, 3/4" across), with blooming flowers at the tip and fruits maturing below. Individual flowers are about 1/8" across, with a narrow, very hairy, basal calyx extending into five sharply pointed lobes, and a narrow, tubular corolla that abruptly flares into five rounded lobes ("petals") surrounding a white or greenish-white throat; the upper two lobes are slightly smaller than the lower three. Both the calyx and corolla are the same color whether light blue, purple, or pink. Hidden in the throat of the corolla are four stamens with yellow anthers and a short style. The fruit consists of a capsule containing four nutlets that is nestled in the remains of the calyx. This plant is a pioneer in highly degraded habitats that is especially fond of cracks and crevices — despite its weedy appearance, prostrate vervain is a native species. In spite of the tiny size of its anatomical characters, this is a relatively straightforward plant to confidently identify from the hairy stems, leaves, and flower spikes and the tiny, lavender apical flowers.

Verbena

Verbena bracteata

prostrate vervain [Blooms: Jun-Aug]

Flower spikes about 5" long cover the top of the blue vervain plants; they are densely covered with floral buds and flowers that bloom from the bottom of the spikes sequentially to the top. Below the open flowers are maturing fruits. Individual flowers are tubular, 1/4" long and across, with a grayish green to dark red, hairy calyx sporting five narrow teeth; a tubular, purplish-blue corolla twice the length of the calyx that abruptly flares into five equal, ovate lobes ("petals"); four recessed stamens; and a single, short style. The throat of the flower is "bearded" (occluded by light blue filaments extending to the middle of the opening) which hides the reproductive organs. This is the only vervain with a blue (not deep violet/lavender) color; other species are pink, white, or lavender. The fruit consists of four 2 mm long nutlets held in the remnants of the calyx. Blue vervain might be confused with hoary vervain, but the latter has larger (2x), lavender (not purplish-blue) flowers (with one lobe/petal notched and two larger than the other three), and ovate to egg-shaped (not lance-shaped) leaves that are sessile (not on petioles 1/2" long or longer). Blue vervain is especially common along the margins of the lagoons of Jackson Park.

Verbena

Verbena hastata

blue vervain [Blooms: Jul-Sep]

Hoary vervain flower spikes are 1-8" long; they are crowded with buds and bluish violet to lavender flowers 1/2" across that open sequentially from the bottom of the spike to the top; below the open flowers are maturing fruits. Individual flowers are tubular, 1/2" long and across, with a grayish green to reddish-purple, hairy calyx sporting five narrow teeth. The tubular, bluish violet to lavender corolla, twice the length of the calyx, abruptly flares into five ovate lobes ("petals"); the two lateral lobes are the largest, while the lower lobe(s) has/have a notched tip. Hidden in the tubular corolla are four recessed stamens with yellow anthers and a single, short green style. The fruits consist of four nutlets held in the remnants of the calyx.

Verbena

Verbena stricta

hoary vervain [Blooms: Jun-Jul]

White vervain produces numerous 6" long, widely spaced flower spikes, each covered by tiny (1/8" wide) white flowers sparsely distributed along the length of the flower spike. Individual flowers consist of a hairy, tubular green calyx with five lance-like teeth and a curved, tubular corolla that abruptly flares into five short, rounded, brilliant-white lobes ("petals"). The throat of the flower is "bearded" (occluded by white filaments extending to the middle of the opening), hiding four stamens with greenish anthers, a green style, and a two-lobed stigma. The upper (more apical) two lobes of the corolla are slightly smaller than the other three (like prostrate vervain) and the tip of the lowermost lobe of the corolla is notched (like hoary vervain), but you're not likely to confuse white vervain with any other Verbena. The fruit is four oblong nutlets nestled in the remnants of the calyx.

Verbena

Verbena urticifolia

white vervain [Blooms: Jun-Sep]

Thyme-leafed speedwell's upper stems host clusters (racemes) of flowers 2-7" long; flowers may either be opposite or alternate along the stem. Each flower is 1/8-1/4" across with a short (1/4"), slender petiole; at the base of the petiole is an oblong, hairless, leafy bract also about 1/4" long with smooth margins. (Thus, the flowers effectively arise from leaf axils.) Individual flowers consist of a green calyx with four triangular lobes shorter than the petals; a short corolla with four oval petal-like, lobes (the lowest one smaller than the other three); two stamens with purple or white anthers; and a green ovary with a single style. The lobes (petals) are white or pale blue, usually with dark blue lines radiating from the flower's center on the upper three lobes. The fruit is a laterally flattened, heart-shaped capsule 1/4" across with two chambers and a fringe of glandular hairs. When mature, the chambers split open, releasing 50-72 tiny seeds. There are 16 species of the genus Veronica reported from Illinois. I have only seen three (Veronica arvensis, Veronica polita, and Veronica serpyllifolia) in Jackson Park but, given their minute size and inconspicuousness, it seems likely that more remain to be found.

Veronica

Veronica serpyllifolia

thyme-leafed speedwell [Blooms: Apr-?]