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Northern evening primrose flowers are 3/4-1.5" across, have four yellow, heart-shaped petals, eight yellow stamens, and a cross-shaped stigma on a modest style; the stamens are as long or longer than the petals and the stigma is nearly fills the floral opening. At the base of the flower are four sepals 1/2-1" long, that bend back along the 3/4-1.5" long floral tube as the flower develops with pairs of sepals attached along one edge until the flower opens. Only northern evening primrose with a distinct ridge or knob near the tips of the sepals. The seed capsule develops from the ovary; it is 3/4-1.5" long, cylindrical but tapering slightly towards the tip, with four robust, flaring lobes at the tip. — — — There are (at least) three evening primroses in Jackson Park — common evening primrose (Oenothera biennis), northern evening primrose (Oenothera parviflora), and hairy evening primrose (Oenothera villosa). They are easily confused with each other. (Judging by the conflicts between the accounts on Illinois Wildflowers [https://www.illinoiswildflowers.info/], Minnesota Wildflowers [https://www.minnesotawildflowers.info/], and Missouri Plants [http://www.missouriplants.com/], even the experts are confused.) All three species have yellow flowers with an X-shaped stigma, reflected sepals, petals with a small notch in their apex, stems that may be red or green, and lance-shaped leaves with short, widely-spaced teeth. Northern evening primrose is distinguished by (1) the presence of a knob or ridge at the tip of each sepal which the other two species lack (2) a seed capsule with four robust, flaring lobes with rounded tips at the apex of the seed capsule, and (3) a stigma that is very robust and almost fills the opening of the flower. Northern evening primrose has green to yellowish-green sepals and is the only one of the three evening primroses to (sometimes) have reddish color on some or all of the midvein of the leaves. Hairy evening primrose (often but not always) has (1) striped or reddish-tinged sepals (check the buds where this character is often easier to see), (2) flowers that turn orange as they begin to senesce (although the reliability of this character is uncertain), and (3) always has glandular hairs covering the ovary that have "pimple-like" bright-red bases; these red pimples may also occur on the stems. All three species are somewhat similar to prairie sundrop (O. pilosella); the latter has very hairy stems and leaves, and translucent lines that radiate from the base of the petals in the flowers.

Oenothera

Oenothera parviflora

northern evening primrose [Blooms: Jun-Jul]

Prairie sundrop is similar to evening primrose (O. biennis) but more delicate and only about 2' tall. The flowers are ~2" across, pale golden-yellow, with translucent lines that radiate from the petals' bases. The flower is attached to a 1/2-1" long floral tube whose base is the ovary, nestled in a leaf axil. The floral tube is covered by four hairy sepals; the four petals are heart-shaped; the stamens are long, yellow, with large yellow anthers; the style extends well beyond the stamens and has a large, X-shaped stigma. Each flower lasts only about a day. The fruits form from the ovary which often have four prominat wings. The fruits are ~1/2" long, four-angled or ridged, club-shaped, with four internal chambers filled with 1 mm long seeds.

Oenothera

Oenothera pilosella

prairie sundrop [Blooms: Jun]

Hairy evening primrose produves an elongating spike of flowers arises from the apex of the stem; flowers bloom near the tip while older flowers and seed capsules develop below. Individual flowers are 3/4-1.5" across; they sit on a long floral tube with the ovary at its base (attached to a leaf axil). The flowers have four yellow, heart-shaped petals, eight yellow stamens, and a cross-shaped stigma on a modest style; at the base of the flower are four sepals 3/8-3/4" long, reflected back along the floral tube as the flower develops with pairs of sepals attached along one edge until the flower opens. The sepals are often red-striped or red-tinged. Note the long, hairy floral tube below the sepals, the reddish stem, and glandular hairs with red bases on the ovary. The seed capsule develops from the ovary; it is up to 1.75" long, cylindrical but tapering slightly towards the tip, with four tiny lobes at the tip. — — — There are (at least) three evening primroses in Jackson Park — common evening primrose (Oenothera biennis), northern evening primrose (Oenothera parviflora), and hairy evening primrose (Oenothera villosa). They are easily confused with each other. (Judging by the conflicts between the accounts on Illinois Wildflowers [https://www.illinoiswildflowers.info/], Minnesota Wildflowers [https://www.minnesotawildflowers.info/], and Missouri Plants [http://www.missouriplants.com/], even the experts are confused.) All three species have yellow flowers with an X-shaped stigma, reflected sepals, petals with a small notch in their apex, stems that may be red or green, and lance-shaped leaves with short, widely-spaced teeth. Northern evening primrose is distinguished by (1) the presence of a knob or ridge at the tip of each sepal which the other two species lack (2) a seed capsule with four robust, flaring lobes with rounded tips at the apex of the seed capsule, and (3) a stigma that is very robust and almost fills the opening of the flower. Northern evening primrose has green to yellowish-green sepals. Hairy evening primrose (often but not always) has (1) striped or reddish-tinged sepals (check the buds where this character is often easier to see), (2) flowers that turn orange as they begin to senesce (although the reliability of this character is uncertain), and (3) always has glandular hairs covering the ovary that have "pimple-like" bright-red bases; these red pimples may also occur on the stems. All three species are somewhat similar to prairie sundrop (O. pilosella); the latter has very hairy stems and leaves, and translucent lines that radiate from the base of the petals in the flowers.

Oenothera

Oenothera villosa

hairy evening primrose [Blooms: Aug-Oct]

Lady's-thumb is an exotic, native to Europe, but has a long history in North America — it was first reported from the Great Lakes in 1843. Flower spikes (racemes) are 1/2-1.5" long and densely covered with tiny (1/8" long) pinkish-white flowers; flower color is quite variable, even within a single flower spike. The flowers are "shy" and often open only at the tips of the 4 or 5 sepals (there are no petals); both the six stamens and the single style (divided into three towards the distal end) remain hidden. Seeds are black; they develop singly within the persistent sepals and remain largely hidden by the sepals; they can be found on flower heads still blooming. Lady's-thumb prefers wet habitats.

Persicaria

Persicaria maculosa

lady's-thumb [Blooms: Jun-Sep]

A relative of the smartweeds and lady's thumb, jumpseed consist of a single, unbranched stem 1-2' tall with an apical inflorescence that can add an additional 16". Flowers are brilliant white, tiny (<3/16" across), with 4 sharply pointed petals (actually, sepals); 4-5 white stamens and a style are visible if you have really good eyesight. Flowers are widely spaced along the flower stalk on very short green stalks. Jumpseed is easy to tell from other smartweeds by the very long inflorescence with sparsely distributed flowers and very large leaves.

Persicaria

Persicaria virginiana

jumpseed [Blooms: Jul-Aug]

Sweet mock-orange inflorescences are racemes with 5-7 fragrant flowers; the most basal pair of flowers arise directly from the leaf axils. Individual flowers are 1-1.5" across, have four triangular green sepals, four brilliant white petals, 20-50 white stamens with orange anthers, and four white styles with yellow-orange stigmas. The styles are closely appressed proximally, separate distally.

Philadelphus

Philadelphus coronarius

sweet mock-orange [Blooms: May-Jun]

Common plantain is an introduced plant commonly found in lawns and disturbed areas. The flower stalks are erect, 4-20" long; the lower third is green and unadorned while the upper two-thirds is densely covered with 1/8" green flowers. Individual flowers consist of four green sepals with a green keel and membranous margins above a single green bract, four filamentous stamens with tan to purplish anthers, a single white style, and a four small, papery, petals. The flowers are wind-pollinated. The fruits are rounded, egg-shaped capsules about 1/8" long. Common plantain is nearly indistinguishable from (the native) black-seeded plantain (Plantago rugelii); the two species differ only in the details of the sepals and seed pods. (Common plantain's seed capsule is egg-shaped and splits around the middle; black-seeded plantain's seed capsule are 2-3 times longer than wide splits nearer the bottom.) One, more ephemeral, character may help distinguish the two species in the field. According to Minnesota Wildflowers, the base of the leaf stalks of the native species (Rugel's plantain, P. rugelii) are dark red or purple in color; the base of the leaf stalks of the exotic common plantain (P. major) are uniformly green. Rugel's plantain is probably as common and tolerant of a range of habitats as common plantain, but specimens of this plant tend to be reflexively referred to "common plantain."

Plantago

Plantago major

common plantain [?-Aug]

In late spring and early summer, curly pondweed produces both flowers and turions (a detachable, dormant bud, a.k.a. a "winter bud"). Cylindrical flower spikes 1-1.5" long arise from the tips of the stems or the axils of upper leaves; peduncles up to 3" long hold the flower spikes just above the water. Each flower spike may have 3-5 whorls of flowers. Individual flowers are tiny (3 mm across); they consist of four greenish-brown or greenish-red sepals, four stamens with a ladle-shaped appendage, and four ovules topped by a four-part style. Flowers may be pollinated by either wind or water. Fertilized ovules produce a dry seed (a maximum of four per flower) that is generally oval but asymmetrical in detail with a knobby keel along one edge and a long, curved beak or tooth at the opposite end. Seed germination has never been observed. Simultaneously, the plant produces turions — rosettes of small, holly-like leaves that overall have a cone-like appearance — from the tips of stems and leaf axils. After the fruits and the turions are released, the rest of the plant degenerates over the course of the summer. The turions sink to the bottom of the water body and root, forming small plants that overwinter.

Potamogeton

Potamogeton crispus

curly pondweed [Blooms: May-?]

Upright prairie coneflower (aka, "Mexican hat") is a 1-3' tall plant native to North America but an exotic in Illinois; it looks like English plantain (Plantago lanceolata) dressed up for a party. The stems are green and longitudinally ridged; they bear alternate, pinnatifid leaves with 5-11 elongate, narrow lobes. The upper stems produce flower stalks (peduncles) 2-12" long, minutely grooved, and lacking leaves; the peduncles are very "wire-like." The flowerheads are 1.5-3" long cylinders composed of numerous brown or purplish disk florets that bloom from the bottom of the cylinder to the top; at the base of the cylinder are 4-12 ray florets with drooping, 1/2-1.5" long, oblong petals with notched tips that are yellow, maroon, or yellow with basal maroon patches (the "party skirt"). This plant is unmistakable when in bloom. The imaged flower was photographed in Bobolink Meadow in July, 2010; I have not seen a specimen since.

Ratibida

Ratibida columnifera

upright prairie coneflower [Blooms: Jul]

In buckthorn, umbels of flowers arise from leaf axils. Flowers on a single plant may be perfect (both male and female organs present), but more commonly the flowers are dioecious with either all male (staminate) or all female (pistillate) flowers found on any given plant. Individual flowers are 1/4" across. They have four greenish-yellow, triangular, spreading sepals that together form a calyx and four petals that can't be easily seen without magnification; the "petals" in the flowers are actually sepals. Male flowers have four greenish stamens with cream-colored anthers; female flowers have a single style that has a tip divided into four branches. After fertilization, female (or perfect) flowers produce a shiny black, 1/4" wide, cherry-like fruit containing 3-4 seeds. Birds can eat the fruit without ill effects, but in humans the fruit produces stomach cramps and acts as a strong laxative (as indicated by the species name, "cathartica").

Rhamnus

Rhamnus cathartica

buckthorn [Blooms: May-Jun]

Jetbead flowers are solitary, arising on the tips of the green twigs. Flowers are elevated on 1/4-1/2" long stalks. There are four green sepals at the base of the flower, each with a bractlet half the length of the sepals that is located below the intersection of each pair of sepals. Four white, nearly circular petals alternate in position with the sepals. A greenish hypanthium covers the center of the flower with a four-lobed ridge marking its edges. Numerous (30-60) stamens with white filaments and yellow anthers, usually in four distinct clusters, lie inside the ridge; the stamens surround four pistils, each with a single style and a circular, terminal stigma. After fertilization, 1-4 fruits develop in the center of the flower that turn shiny black at maturity (the "jetbead"); the seeds remain attached to the persistant sepals until late in the fall and may persist through the winter.

Rhodotypos

Rhodotypos scandens

jetbead [Blooms: Apr-May]

The upper stems of marsh yellow-cress end in one or more 2-8" long racemes of flowers; racemes may also develop from leaf axils. Flowers in bloom are restricted to the tip of the raceme; cylindrical seedpods (siliques) oriented perpendicular to the stem decorate the lower regions of the raceme. Individual flowers are tiny (about 3 mm long) with four greenish-yellow oblong sepals, four yellow spatulate petals, 4-6 stamens with yellow or light brown anthers, and a stout style with a two-lobed stigma. After fertilization, 3-10 mm long, dumbbell-shaped, hairless, green siliques containing two rows of tiny seeds develop. The siliques bear the remnant of the style on one end and 1/2" long pedicels on the other.

Rorippa

Rorippa palustris

marsh yellow-cress [Blooms: Jun-Aug]

Sweet coneflower flowerheads are 2-3.5" across, occurring as single blooms on the tips of stems or branches. A flowerhead is comprised of numerous 1/2" long medium green, lance-shaped phyllaries, downy with short hairs; 10-20 bright yellow, oblong, sterile ray florets with a tiny notch at their tips; and a deep brown hemisphere of 200-400 disk florets (the central disc). The disk florets have a tubular, dark reddish-purple corolla; bifurcated stigmas that have sharp tips; with the purplish-brown anthers adhering side by side around the deep purple styles. Sweet coneflower is distinctive among the Jackson Park coneflowers with its chocolate brown disk and bright yellow, elongate-oval petals with a shallow notch at their tips.

Rudbeckia

Rudbeckia subtomentosa

sweet coneflower [Blooms: Jul-Aug]

Wild mustard's upper stems end in racemes of 1/2" wide yellow flowers. Each flower has four yellow petals, six yellow stamens, and a single green pistil. After fertilization, the pistil elongates into a thin, cylindrical silique (seedpod), 1/2-1.5" long, filled with tiny (2 mm diameter) brown or black seeds; the distal third of the silique is a ridged or square "beak". The seeds are reported to remain dormant but viable in the soil for up to 60 years.

Sinapis

Sinapis arvensis

wild mustard [Blooms: May-Jun]

Hedge mustard flowers occur at the tips of long, slender flower stalks that curve upward; the flower stalks typically arise in abundance at the top of the plant. Each flower is small (1/4" across) with four greenish-yellow sepals, four rectangular yellow petals with rounded tips, a single stout style, and six stamens. The fruit is a very slender, cylindrical seedpod (silique) lower on the flower stalk than the blooming flowers; the siliques are 3/8-3/4" long, erect, oriented parallel to and often appressed to the flower stalk, with the remains of the style on their tips. The siliques turn light brown when mature and split open to release the seeds. The short (<3/4"), straight, appressed seedpods and the upward-curving, sparse flower stalks with only a few blooms open at any one time distinguish hedge mustard from other local mustards.

Sisymbrium

Sisymbrium officinale

hedge mustard [Blooms: May-Jul]

The axils of the middle and upper leaves of woodland goldenrod produce small clusters of 1-12 flowerheads; the stem tip may also develop a small (3" long, 1.5" across) panicle of flowerheads. Individual flowerheads are tiny (3 mm) consisting of 4-5 ray florets with yellow, oblong-elliptical "petals" and 3-6 disc florets with golden yellow, tubular corollas with five tiny lobes. At the base of each flowerhead are three overlapping series of light green, elongate-elliptical bracts (phyllaries). The fruit is a small, bullet-shaped seed 1-2 mm long with a tuft of white hairs 3-5 mm long. In Illinois, only woodland goldenrod and broad-leafed goldenrod (Solidago flexicaulis) produce clusters of flowerheads from the axils of the middle and upper leaves; however, the latter has coarsely serrated leaves with petioles and should be easy to distinguish from woodland goldenrod.

Solidago

Solidago caesia

woodland goldenrod [Blooms: Sep-Oct]

The tip of broad-leaved goldenrod's stem produces a cylindrical inflorescence (a raceme or a narrow panicle) 1.5-5" long and about a third as wide; smaller racemes (1/2-1.5" long, smaller than the leaves) arise from the axils of the upper leaves. Individual flowerheads are about 1/4" across with 3-4 yellow ray florets asymmetrically deployed around the flowerhead and 4-8 yellow disk florets. The base of the flowerhead is surrounded by 3-5 series of unequal, narrowly oblong, light green phyllaries; the outermost phyllaries are 1-2 mm long while the innermost are 3.7-4.7 mm long. Broad-leaved goldenrod prefers high-quality, moist, shaded woodlands. In Illinois, only broad-leaved goldenrod and woodland goldenrod (Solidago caesia) produce clusters of flowerheads from the axils of the middle and upper leaves. Broad-leaved goldenrod has unusually large flowers for a goldenrod and the ray florets are not evenly distributed around the flowerhead. In addition, woodland goldenrod has smooth or finely serrated leaf margins (rather than coarsely-toothed leaves) and no petioles (the leaves are sessile) — it should be easy to distinguish from broad-leaved goldenrod.

Solidago

Solidago flexicaulis

broad-leaved goldenrod [Blooms: Jul-Sep]

Early goldenrod's inflorescence is apical, a panicle of flowering stems that "often arch upward and outward like a fireworks display" (John Hilty, www.illinoiswildflowers.info). Individual flowerheads are about 1/4" across; they have 4-12 yellow ray florets, not all open at one time, and 8-15 yellow disk florets. The base of the flowerhead is surrounded by 3-4 series of overlapping, narrow, hairless, yellowish-green bracts (phyllaries) that together are 3-4 mm (1/16") long. The pedicels (flower stalks) are usually hairless, all attached on one side of the panicle's flowering branches and curving upward. The fruit is a cone-shaped dry seed 1-1.5 mm long with an attached tuft of light brown hairs 2.5-3.5 mm long. The combination of wing-like leaflets in the leaf axils, hairless stems and leaves, and an inflorescence whose flowering stems flare up and out is diagnostic for early goldenrod. Field goldenrod (Solidago nemoralis) is much the same size and general appearance, but has hairy stems and leaves, unlike early goldenrod.

Solidago

Solidago juncea

early goldenrod [Blooms: Jul/Aug-?]