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Sweet mock-orange inflorescences are racemes with 5-7 fragrant flowers; the most basal pair of flowers arise directly from the leaf axils. Individual flowers are 1-1.5" across, have four triangular green sepals, four brilliant white petals, 20-50 white stamens with orange anthers, and four white styles with yellow-orange stigmas. The styles are closely appressed proximally, separate distally.

Philadelphus

Philadelphus coronarius

sweet mock-orange [Blooms: May-Jun]

Cleft phlox flowers are supported by 1" pedicels, are 3/4" across, white or pale violet, with a tubular corolla that flares into five petal-like lobes that are each deeply cleft at their tips; the throat is very narrow. A three-lobed stigma and the orange anthers of the stamens lie just within the opening to the throat of the flower. Cleft phlox is endemic to the Midwest. The cleft petal lobes distinguish cleft phlox from all other Phlox species.

Phlox

Phlox bifida

cleft phlox [Blooms: Apr-?]

Woodland phlox is a 1-1.5' tall plant with a fuzzy to hairy stem, light green or reddish brown in color. Leaves are opposite (or sometimes in a three-leaf whorl), sessile or clasping the stem, three times longer than wide, lance-shaped with smooth margins, and fuzzy to hairy overall with short hairs around the margins. The flowers are 1" across, pale blue-violet, lavender, or (rarely) white, and with a long, narrow tubular throat that abruptly flares into five petal-like, spoon-shaped lobes. The calyx is green or reddish-green, hairy, with five long, narrow teeth almost half the length of the flower, spreading near their tips. The five stamens and style are hidden within the throat. Woodland phlox has relatively broader leaves than other Phlox species and has a more intense blue-violet (and less pink) color.

Phlox

Phlox divaricata

woodland phlox [Blooms: Apr-May]

Smooth phlox is a 1-2' tall plant with an unbranched, hairless stem topped with a slightly domed cluster of 3-20 pink or lavender flowers. Each flower is about 3/4" across and has a long, tubular throat that abruptly flares into five petal-like lobes; each lobe is wedge-shaped and overlaps the next lobe clockwise. The calyx (also hairless) consists of five triangular segments that taper rapidly to sharp points; the calyx is <1/3 the length of the corolla tube. The five stamens and single style with a three-part stigma either lie within the corolla tube or are just visible in the throat. After fertilization, a narrow seed capsule filled with numerous seeds fills the persistant calyx. Smooth phlox prefers moist to wet habitats. Smooth phlox flowers resemble prairie phlox (P. bifida) but the latter species is quite hairy on the stems, leaves, and the exterior of the flower (the calyx); smooth phlox is hairless. Checking the hairiness (or lack thereof) of the calyx is the quickest way to tell prairie phlox from smooth phlox.

Phlox

Phlox glaberrima interior

smooth phlox [Blooms: Jun-Aug]

Prairie phlox is a compact, single-stemmed, plant 1/2-1.75' tall with (typically) a dome-shaped cluster of flowers at the apex. Individual flowers are 1/2-3/4" across, pink or lavender (rarely white); the corolla is tubular with five wedge-shaped, petal-like lobes often with darker markings (nectar guides) near the throat; the corolla is hairy on all external surfaces. The throat is very narrow; the five stamens with yellow anthers are recessed but just visible. The very short, tubular, hairy calyx base is green, with five long, narrow teeth that extend parallel to the narrow throat (also hairy) of the corolla. The fruit is oval and shorter than the calyx (in which it develops); when ripe, the capsule splits explosively, flinging the seeds in all directions. The species name ("pilosa") means "hairy" in Latin, and this species is indeed hairy on almost every surface. Prairie phlox flowers resemble smooth phlox (P. glaberrima) and, to a lesser extent, woodland phlox (P. divaricata), but the former species is quite hairy on the stems, leaves, and the exterior of the flower (the calyx); smooth phlox is hairless. Checking the hairiness (or lack thereof) of the calyx is the quickest way to tell prairie phlox from smooth phlox; checking the hairiness of the exterior of the corolla tube will distinguish woodland phlox (hairless) from prairie phlox (hairy).

Phlox

Phlox pilosa

prairie phlox [Blooms: Apr-Jun]

Obedient plant flower spikes are up to 10" long, and have four rows of horizontal, tubular flowers or flower buds, with a stem densely covered with fine hairs (usually invisible beneath the tightly packed flowers). Individual flowers are purplish-pink, lavender, or white, with darker spots or stripes on the interior. Each 1" long flower consists of a hood-like upper lip and a broad lower lip with two small lateral lobes and a larger medial lobe. Four stamens with purple anthers lie just below (but within) the upper lip. The style lies among the stamens but is slightly longer; it bears a bifurcated stigma. The ~1/2" long calyx is green, tubular, with five sharp, flaring triangular lobes; it is densely covered with short, fine hairs. The fruit consists of four nutlets housed in the persistant calyx. Obedient plant will grow in most sunny habitats but prefers moist sites near water.

Physostegia

Physostegia virginiana

obedient plant [Blooms: Jul-Oct]

Mayapple produces a single, hairless stem that branches no more than once; each stem thus has either a single leaf (infertile stems) or two leaves (fertile stems). Fertile stems produce a solitary nodding flower, 2-3" across, that is hidden beneath the leaves where the two leaves branch on the stem. The flowers contain 6-9 brilliant white petals and a ring of 12-18 tan stamens with large, banana-shaped anthers, all surrounding a surprisingly large, yellow ovary topped with a button of a stigmata. The fruit is a large (1.5" long), ellipsoidal berry, yellow when ripe. All parts of the plant are poisonous including the green fruit, but the ripe (yellow) fruit is edible (in small quantities).

Podophyllum

Podophyllum peltatum

mayapple [Blooms: Apr-May]

Heal-all (a.k.a., self-heal) is a native species superficially similar to ground ivy (Glechoma hederacea) or early-season henbit (Lamium amplexicaule). Heal-all has the characteristics of other low-growing plants in the mint family — an upright, four-angled stem, with lines of hairs along the ridges, 1' tall or less, with a short (1-2") spike of flowers at the apex — and is thus easy to overlook. But the tiny (1/2" long), tubular, two lipped flowers are distinctive (and charming). The light purple or blue upper lip is bent at right angles to form a hood over the aperture with the four stamens and single style with a bifurcated stigma at its tip curving along inside of the hood. The lower lip is divided into three lobes, the middle one white and fringed; the two smaller lateral lobes are the same color as the hood. The hood of the upper lip also sports a "mohawk" haircut of relatively long hairs along the crest of the hood. The flower calyx is light green or reddish, hairy along the edges. After fertilization, four tiny, brown, finely-ridged nutlets develop within the persistant calyx; the seeds are reported to be dispersed by a mechanism that uses the energy in raindrops impacting the calyx to fling the seeds away from the mother plant. The leaves are opposite, up to 2" long and half as wide with a short petiole; they are egg- or lance-shaped with a central vein that is hairy on the underside. Heal-all prefers moist habitats with a history of disturbance.

Prunella

Prunella vulgaris

heal-all [Blooms: Jun-Aug]

Slender mountain-mint flowers occur in small terminal clusters at the tips of the upper stems with distinctive very long, narrow bracts with spine-like tips just beneath the clusters; the bracts are densely covered with short, curled hairs on the blade and longer, bristly hairs along the margins. Individual flowers are tubular, white with scattered purple dots, and about 1/4" long; there is a calyx that is divided several slender green lobes. The corolla is divided into two parts, an upper lip and a lower lip that is divided into three lobes. The four stamens and two styles are also white, but the anthers are purple. The styles have a bifurcated stigma. Slender mountain mint is very similar to Virginia mountain mint (Pycnanthemum virginianum), but several characters make it straightforward to distinguish them. Leaves on slender mountain mints are always less than 1/4" wide and the stems are hairless; Virginia mountain mint will have the larger leaves more than 1/4" wide and has lines of white hairs on the stems. Slender mountain blooms earlier than Virginian mountain mint, but the blooming period of the two species overlaps. All parts of slender mountain mint give off a strong mint-like odor when crushed, but please restrain yourself from excessive foliage-crushing — it's not polite.

Pycnanthemum

Pycnanthemum tenuifolium

slender mountain-mint [Blooms: Jun-Aug]

Virginia mountain-mint flower heads are 3/4-1" across with up to 50 flowers; only a few flowers open at any one time. The flowers are hairy on the outer surfaces, white to light purple in color. Individual flowers are 1/4" across, tubular, and two-lipped; the lower lip is three-lobed with purple spots; four stamens with purple anthers emerge from the lower tube. There are two styles with deeply-notched tips that extend well beyond the edges of the lips of the corolla. The calyx that cups the flower is hairy, light green and has five triangular teeth.

Pycnanthemum

Pycnanthemum virginianum

Virginia mountain-mint [Blooms: Jul-Aug]

Lesser celadine flowers arise singly from the axils of the upper leaves or on the ends of stems. Individual flowers are about 1" across with three light green sepals half the length of the petals, 7-12 yellow, oblong petals, and a ring of about 30 (yellow) stamens around a central group of about 15-20 (greenish) pistils. This species rarely produces fruit, instead reproducing asexually. The alternative name "Ficaria verna" is occasionally used for lesser celadine, but that name is now considered invalid.

Ranunculus

Ranunculus ficaria

lesser celadine [Blooms: Apr-May]

In buckthorn, umbels of flowers arise from leaf axils. Flowers on a single plant may be perfect (both male and female organs present), but more commonly the flowers are dioecious with either all male (staminate) or all female (pistillate) flowers found on any given plant. Individual flowers are 1/4" across. They have four greenish-yellow, triangular, spreading sepals that together form a calyx and four petals that can't be easily seen without magnification; the "petals" in the flowers are actually sepals. Male flowers have four greenish stamens with cream-colored anthers; female flowers have a single style that has a tip divided into four branches. After fertilization, female (or perfect) flowers produce a shiny black, 1/4" wide, cherry-like fruit containing 3-4 seeds. Birds can eat the fruit without ill effects, but in humans the fruit produces stomach cramps and acts as a strong laxative (as indicated by the species name, "cathartica").

Rhamnus

Rhamnus cathartica

buckthorn [Blooms: May-Jun]

Jetbead flowers are solitary, arising on the tips of the green twigs. Flowers are elevated on 1/4-1/2" long stalks. There are four green sepals at the base of the flower, each with a bractlet half the length of the sepals that is located below the intersection of each pair of sepals. Four white, nearly circular petals alternate in position with the sepals. A greenish hypanthium covers the center of the flower with a four-lobed ridge marking its edges. Numerous (30-60) stamens with white filaments and yellow anthers, usually in four distinct clusters, lie inside the ridge; the stamens surround four pistils, each with a single style and a circular, terminal stigma. After fertilization, 1-4 fruits develop in the center of the flower that turn shiny black at maturity (the "jetbead"); the seeds remain attached to the persistant sepals until late in the fall and may persist through the winter.

Rhodotypos

Rhodotypos scandens

jetbead [Blooms: Apr-May]

Hairy wild petunia is, like the name says, hairy — stems, leaves, and petioles all densely covered with long (1-2 mm) white hairs on all surfaces. The plant is short, generally 8-12" tall. The leaves are opposite, twice as long as wide, oval or triangular, with smooth margins and with short (1-3 mm) hairy petioles or sessile. The flowers emerge directly from upper leaf axils; what looks like a flower stalk is actually part of the corolla tube. The flowers are pale lavender or pale purple, about 2" across, funnel shaped with five lobes tapering rapidly to a narrow throat, four white to purplish stamens, and a white to purplish style with an asymmetrically split stigma. There are purple streaks (nectar guides) on the lowermost lobe's throat region. The short, tubular calyx is green, with five long, narrow teeth that extend for half the length of the flower; the calyx, too, is hairy. The fruit is a bullet-shaped capsule, shorter than the calyx teeth (among which it nestles); the seeds are covered with fine spiraling hairs that allow the seeds to self-bury. The flowers usually last only one day. Hairy wild petunia is a distinctive plant unlikely to be confused with any other.

Ruellia

Ruellia humilis

hairy wild petunia [Blooms: Jun-Sep]

Upper branches of elderberry end in flat to domed panicles of flowers 3-10" across. Individual flowers are about 1/4" across, with a green, cup-shaped calyx, a creamy white corolla with five petal-like lobes, five long, spreading stamens with white anthers, and a creamy white hemispherical ovary with 3-5 short styles; flowers occur in umbel-like panicles 10" across made of dozens of flowers. The fruits are <1/4" wide spherical drupes in starburst-like sprays, dark purple to black when mature. The flower stalks are light green, turning purplish as the fruit ripens. The raw fruit is sweet with a slightly bitter or astringent aftertaste but may induce nausea or diarrhea if eaten in more than small quantities. The fruit is widely used in a variety of herbal supplements (as infusions and extracts), jam, and herbal wine. Note that all parts of the plant including the sap (except the flesh of the fruit) contain toxic cyanide-producing glycosides; berries should be cooked before consumption.

Sambucus

Sambucus nigra

elderberry [Blooms: May-Jul]

Soapwort produces clusters of 3/4-1" flowers from the tips of terminal stems and branches. Individual flowers have a 1" long, green or reddish, sometimes hairy, cylindrical calyx; a corolla with five spreading (later reflexed), pink to white, oblong petals/lobes with a notched tip; 10 white stamens with pale yellow anthers that protrude from the throat of the flower; and a similarly protruding, divided white style. The calyx persists as an inflated seed capsule filled with 15-75 tiny dark brown, kidney-shaped seeds. Soapwort might be mistaken for a phlox, but the stamens and style in phlox are always hidden in the calyx tube, never protruding as in soapwort, and soapwort has leaves with three obvious longitudinal veins. Soapwort contains saponins in both the foliage and (especially) the seeds that make a frothy soap-like lather when boiled (and were once used as a soap substitute), but caution is recommended — the saponins are toxic. Herbivores generally avoid this plant.

Saponaria

Saponaria officinalis

soapwort [Blooms: Jun-Jul]

Early figwort (aka, lanceleaf figwort) is difficult to distinguish from late- (or Maryland-) figwort (S. marilandica) except by blooming date (before July = early, after July = late, July = could be either) and details of the flowers (especially the sterile stamen). The inflorescence (a panicle) is constructed from short, spreading, sparse branches covered in glandular hairs, bearing a 3/8" long, shiny tubular flower whose outside surfaces are reddish-purple to reddish-green and inner surfaces are reddish brown (upper and lateral lobes) or greenish yellow (lower lobe). Four yellow, fertile stamens and a single blunt, slender style hug the floor of the flower, the style protruding further than the fertile stamens. A single sterile stamen (a "staminode") is located along the upper interior of the flower. The sterile stamens are yellow or green with a broad tip in early figwort (S. lanceolata); they are reddish-brown to purple and narrow in late figwort (S. marilandica). The fruit is a teardrop-shaped capsule, dark brown when mature, that splits in two longitudinally to release the numerous tiny seeds.

Scrophularia

Scrophularia lanceolata

early figwort [Blooms: May-Jul]

Late figwort's inflorescence (a panicle) gives rise to short, spreading, sparse branches covered in glandular hairs, bearing a 3/8" long, shiny tubular flower whose outside surfaces are reddish-purple to reddish-green and inner surfaces are reddish brown (upper and lateral lobes) or greenish yellow (lower lobe). Four yellow, fertile stamens and a single blunt, slender style hug the floor of the flower, the style protruding further than the fertile stamens. A single sterile stamen (a "staminode") is located along the upper interior of the flower. The sterile stamens are yellow or green with a broad tip in early figwort (S. lanceolata); they are reddish-brown to purple and narrow in late figwort (S. marilandica). The fruit is a teardrop-shaped capsule, dark brown when mature, that splits in two longitudinally to release the numerous tiny seeds.

Scrophularia

Scrophularia marilandica

late figwort [Blooms: Jul-Aug]