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Lesser celadine flowers arise singly from the axils of the upper leaves or on the ends of stems. Individual flowers are about 1" across with three light green sepals half the length of the petals, 7-12 yellow, oblong petals, and a ring of about 30 (yellow) stamens around a central group of about 15-20 (greenish) pistils. This species rarely produces fruit, instead reproducing asexually. The alternative name "Ficaria verna" is occasionally used for lesser celadine, but that name is now considered invalid.

Ranunculus

Ranunculus ficaria

lesser celadine [Blooms: Apr-May]

In buckthorn, umbels of flowers arise from leaf axils. Flowers on a single plant may be perfect (both male and female organs present), but more commonly the flowers are dioecious with either all male (staminate) or all female (pistillate) flowers found on any given plant. Individual flowers are 1/4" across. They have four greenish-yellow, triangular, spreading sepals that together form a calyx and four petals that can't be easily seen without magnification; the "petals" in the flowers are actually sepals. Male flowers have four greenish stamens with cream-colored anthers; female flowers have a single style that has a tip divided into four branches. After fertilization, female (or perfect) flowers produce a shiny black, 1/4" wide, cherry-like fruit containing 3-4 seeds. Birds can eat the fruit without ill effects, but in humans the fruit produces stomach cramps and acts as a strong laxative (as indicated by the species name, "cathartica").

Rhamnus

Rhamnus cathartica

buckthorn [Blooms: May-Jun]

Jetbead flowers are solitary, arising on the tips of the green twigs. Flowers are elevated on 1/4-1/2" long stalks. There are four green sepals at the base of the flower, each with a bractlet half the length of the sepals that is located below the intersection of each pair of sepals. Four white, nearly circular petals alternate in position with the sepals. A greenish hypanthium covers the center of the flower with a four-lobed ridge marking its edges. Numerous (30-60) stamens with white filaments and yellow anthers, usually in four distinct clusters, lie inside the ridge; the stamens surround four pistils, each with a single style and a circular, terminal stigma. After fertilization, 1-4 fruits develop in the center of the flower that turn shiny black at maturity (the "jetbead"); the seeds remain attached to the persistant sepals until late in the fall and may persist through the winter.

Rhodotypos

Rhodotypos scandens

jetbead [Blooms: Apr-May]

Hairy wild petunia is, like the name says, hairy — stems, leaves, and petioles all densely covered with long (1-2 mm) white hairs on all surfaces. The plant is short, generally 8-12" tall. The leaves are opposite, twice as long as wide, oval or triangular, with smooth margins and with short (1-3 mm) hairy petioles or sessile. The flowers emerge directly from upper leaf axils; what looks like a flower stalk is actually part of the corolla tube. The flowers are pale lavender or pale purple, about 2" across, funnel shaped with five lobes tapering rapidly to a narrow throat, four white to purplish stamens, and a white to purplish style with an asymmetrically split stigma. There are purple streaks (nectar guides) on the lowermost lobe's throat region. The short, tubular calyx is green, with five long, narrow teeth that extend for half the length of the flower; the calyx, too, is hairy. The fruit is a bullet-shaped capsule, shorter than the calyx teeth (among which it nestles); the seeds are covered with fine spiraling hairs that allow the seeds to self-bury. The flowers usually last only one day. Hairy wild petunia is a distinctive plant unlikely to be confused with any other.

Ruellia

Ruellia humilis

hairy wild petunia [Blooms: Jun-Sep]

Upper branches of elderberry end in flat to domed panicles of flowers 3-10" across. Individual flowers are about 1/4" across, with a green, cup-shaped calyx, a creamy white corolla with five petal-like lobes, five long, spreading stamens with white anthers, and a creamy white hemispherical ovary with 3-5 short styles; flowers occur in umbel-like panicles 10" across made of dozens of flowers. The fruits are <1/4" wide spherical drupes in starburst-like sprays, dark purple to black when mature. The flower stalks are light green, turning purplish as the fruit ripens. The raw fruit is sweet with a slightly bitter or astringent aftertaste but may induce nausea or diarrhea if eaten in more than small quantities. The fruit is widely used in a variety of herbal supplements (as infusions and extracts), jam, and herbal wine. Note that all parts of the plant including the sap (except the flesh of the fruit) contain toxic cyanide-producing glycosides; berries should be cooked before consumption.

Sambucus

Sambucus nigra

elderberry [Blooms: May-Jul]

Soapwort produces clusters of 3/4-1" flowers from the tips of terminal stems and branches. Individual flowers have a 1" long, green or reddish, sometimes hairy, cylindrical calyx; a corolla with five spreading (later reflexed), pink to white, oblong petals/lobes with a notched tip; 10 white stamens with pale yellow anthers that protrude from the throat of the flower; and a similarly protruding, divided white style. The calyx persists as an inflated seed capsule filled with 15-75 tiny dark brown, kidney-shaped seeds. Soapwort might be mistaken for a phlox, but the stamens and style in phlox are always hidden in the calyx tube, never protruding as in soapwort, and soapwort has leaves with three obvious longitudinal veins. Soapwort contains saponins in both the foliage and (especially) the seeds that make a frothy soap-like lather when boiled (and were once used as a soap substitute), but caution is recommended — the saponins are toxic. Herbivores generally avoid this plant.

Saponaria

Saponaria officinalis

soapwort [Blooms: Jun-Jul]

Early figwort (aka, lanceleaf figwort) is difficult to distinguish from late- (or Maryland-) figwort (S. marilandica) except by blooming date (before July = early, after July = late, July = could be either) and details of the flowers (especially the sterile stamen). The inflorescence (a panicle) is constructed from short, spreading, sparse branches covered in glandular hairs, bearing a 3/8" long, shiny tubular flower whose outside surfaces are reddish-purple to reddish-green and inner surfaces are reddish brown (upper and lateral lobes) or greenish yellow (lower lobe). Four yellow, fertile stamens and a single blunt, slender style hug the floor of the flower, the style protruding further than the fertile stamens. A single sterile stamen (a "staminode") is located along the upper interior of the flower. The sterile stamens are yellow or green with a broad tip in early figwort (S. lanceolata); they are reddish-brown to purple and narrow in late figwort (S. marilandica). The fruit is a teardrop-shaped capsule, dark brown when mature, that splits in two longitudinally to release the numerous tiny seeds.

Scrophularia

Scrophularia lanceolata

early figwort [Blooms: May-Jul]

Late figwort's inflorescence (a panicle) gives rise to short, spreading, sparse branches covered in glandular hairs, bearing a 3/8" long, shiny tubular flower whose outside surfaces are reddish-purple to reddish-green and inner surfaces are reddish brown (upper and lateral lobes) or greenish yellow (lower lobe). Four yellow, fertile stamens and a single blunt, slender style hug the floor of the flower, the style protruding further than the fertile stamens. A single sterile stamen (a "staminode") is located along the upper interior of the flower. The sterile stamens are yellow or green with a broad tip in early figwort (S. lanceolata); they are reddish-brown to purple and narrow in late figwort (S. marilandica). The fruit is a teardrop-shaped capsule, dark brown when mature, that splits in two longitudinally to release the numerous tiny seeds.

Scrophularia

Scrophularia marilandica

late figwort [Blooms: Jul-Aug]

Mad-dog skullcap flower racemes may arise at the tips of the stems or from the axils of leaves. Each raceme is up to 6" long, containing 6-7 pairs of flowers; each flower has a short, basal leafy bract. Flowers are up to 8 mm long with a short, slender pedicel, a light green tubular calyx with two shallow lobes (the upper lobe with a dish-like projection), and a pale blue, lavender, or white, densely hairy, tubular corolla 5-8 mm long with a short, hood-like, upper lip and a slightly flared, three-lobed lower lip. Hidden inside the upper lip of the corolla are four stamens and a single bifurcated style. After fertilization, the calyx swells into a green capsule with includes the dish-like projection mentioned above, giving the capsule a unique appearance; inside is a cluster of four nutlets. The mouth of the calyx is visible as a thin slit beneath the projection.

Scutellaria

Scutellaria lateriflora

mad-dog skullcap [Blooms: Jul-Aug]

Evening campion is an exotic from Europe that grows to be 2-3' tall. It is nominally a night-blooming species, but I've observed it in bloom throughout the day in shady areas. Male and female plants are separate. The upper stems end in clusters of one or more 1-1.5" long flowers. The "bladder" (the calyx) is covered with a fuzz of fine hairs and has dark green or reddish-purple lines running from pole to pole (10 lines in male flowers, 20 in female). The five petals are heart shaped (with the point towards the center of the flower) and are deeply cleft into two lobes; the petals form a fringe that makes a distinct collar around the opening to the floral tube. Female flowers have five emergent styles (rarely four or six); male flowers have 10 yellow stamens that barely emerge from the collar at the center of the flower. (Other local Silene sp. have three styles.) The fruit is brown, urn-shaped, and has 10 small teeth on the distal end (or five pairs if you prefer). (Five or six teeth total in S. noctiflora, S. stellata, and S. vulgaris.) Evening campion evolved a dioecious life history (separate male and female plants) about 11 million years ago. (Other species in the genus have "perfect" flowers with both male and female organs present and functional in the flowers.) The evolution and genomic history of the sex-determining Y chromosome (XX = female, XY = male) has been of considerable interest for the light it shines on the evolution of Y chromosomes in both plants and animals (Moraga et al. 2025. Science 387:630–636; Akagi et al. 2025. Science 387:637–643).

Silene

Silene latifolia

evening campion [Blooms: May-Aug]

Royal catchfly's bright red flowers are restricted to the upper regions of the plant as clusters arising from the tips of stems. Individual flowers are 3/4-1" across, have an elongate (1"), narrow, cylindrical calyx with 10 longitudinal ridges, five narrow red or scarlet petals that flare from the tip of the calyx, and stamens with gray anthers that protrude well beyond the opening to the calyx. There are three corkscrew-shaped, threadlike styles that develop after the stamens have matured; like the stamens, they protrude well outside the calyx opening. The fruit is a 1/2" long capsule, open at the top, with 6-8 spreading teeth around the lip. Royal catchfly is hummingbird and butterfly pollinated.

Silene

Silene regia

royal catchfly [Blooms: Jul-Aug]

Starry campion's main stem ends in a panicle of flowers up to 8" long and 3-4" across; additional panicles may arise on the lateral stems on robust plants. Flowers occur in groups of 2-3 on the ends of long, slender branches; the flowers rest directly on short (1/4") pedicels. Individual flowers are 3/4" across. Each flower consists of a bell shaped, light green, fuzzy calyx nearly as broad as it is long with five broad, triangular teeth on its free end; five white petals with a narrow base and a "fringe" of 8-12 narrow lobes on their free ends; 10 stamens with slender white filaments that extend well outside the throat; and three white styles. The styles emerge from a large, green, globular ovary in the middle of the flower. After fertilization, the calyx gets broader (nearly triangular in profile) as the ovary expands into a spherical green ball; the fruit is brown, urn-shaped, and has six teeth on the distal end. The flowers tend to close in bright midday light. They are usually pollinated by moths and, to a lesser extent, bumblebees.

Silene

Silene stellata

starry campion [Blooms: Jun-Jul]

Rosinweed produces a panicle of sunflower-like blooms from the top of the stem and the tops of side stems (if any). Individual flowerheads are 2-3" across; they consist of 12-35 yellow, pistillate (female) ray florets and 70-200+ functionally male disc florets. The green bracts beneath the flowerheads are teardrop-shaped, partially overlapping in two or three layers, and sometimes covered with glandular hairs. The petals of the ray florets have two prominent longitudinal grooves and a tiny apical notch. Only the ray florets can produce seeds — a split style protrudes from a short tube at the base of the ray floret. The central disk is initially green but becomes yellow as the disk florets open; they are yellow, tubular, with five triangular lobes at their free end, with a column of brown stamens with the tips of very long, (non-functional) styles protruding from the tubular corolla. The fruit is a flat seed that develops from the ray florets. (The curious arrangement where only the ray florets can produce seeds while the disk florets are functionally male is true throughout the genus Silphium. The opposite is true in the true sunflowers, genus Helianthus.)

Silphium

Silphium integrifolium

rosinweed [Blooms: Jul-Sep]

A cup plant may have 10-30 flowerheads in bloom at one time. Behind the flowerhead are two or three layers of green bracts; the outer layer is egg shaped with a triangular, pointed, recurving tip. Each sunflower-like flowerhead is about 3-4" across; it has 18-40 yellow ray florets (e.g., the petals) and 85-150 yellow disk florets. The ray florets have a bifurcated but functional style protruding from a short yellow tube at the base of the petal; the disc florets are initially green but transform into yellow, tubular corollas with a non-functional style protruding through the fully-functional stamen. (True throughout the genus Silphium. The opposite is true in the sunflowers, genus Helianthus, where the ray florets are functionally male and the disk florets are fertile, functional females.) The cup-like leaves and square stem distinguish cup plants from other prairie wildflowers.

Silphium

Silphium perfoliatum

cup plant [Blooms: Jul-Sep]

Early goldenrod's inflorescence is apical, a panicle of flowering stems that "often arch upward and outward like a fireworks display" (John Hilty, www.illinoiswildflowers.info). Individual flowerheads are about 1/4" across; they have 4-12 yellow ray florets, not all open at one time, and 8-15 yellow disk florets. The base of the flowerhead is surrounded by 3-4 series of overlapping, narrow, hairless, yellowish-green bracts (phyllaries) that together are 3-4 mm (1/16") long. The pedicels (flower stalks) are usually hairless, all attached on one side of the panicle's flowering branches and curving upward. The fruit is a cone-shaped dry seed 1-1.5 mm long with an attached tuft of light brown hairs 2.5-3.5 mm long. The combination of wing-like leaflets in the leaf axils, hairless stems and leaves, and an inflorescence whose flowering stems flare up and out is diagnostic for early goldenrod. Field goldenrod (Solidago nemoralis) is much the same size and general appearance, but has hairy stems and leaves, unlike early goldenrod.

Solidago

Solidago juncea

early goldenrod [Blooms: Jul/Aug-?]

Saltmarsh sand-spurry flowers consist of five elliptical green sepals, five white to pink petals slightly longer than the sepals, 2-5 stamens with bright yellow anthers, and three short (0.7 mm), thread-like styles and three stigmas. In the Great Lakes region, saltmarsh sand-spurry is widely present along roads where salt is used as a deicer. Some sources refer to this plant as Spergularia marina, but ITIS rejects that binomial; Spergularia salina is the accepted name.

Spergularia

Spergularia salina

saltmarsh sand-spurry [Blooms: May-Aug]

Bladdernut produces white, drooping, bell-shaped flowers in clusters of 5-12 from the axils of the leaves. Each flower is pendant on a long stalk, bell-shaped, about 8 mm long and about 6 mm wide. Flowers have five greenish-white sepals overlapping much of the petals, five bright white petals (visible through the gaps between the sepals) that flare at their tips, five stamens with white filaments and deep yellow-orange anthers, and a greenish-white style with a three-lobed stigma extending beyond the mouth of the bell. The flower stalks (pedicels) are about 1.5-2 times the length of the flower. The fruit is a greenish-white to brown, papery pod (like a "Chinese lantern") that is about 3" long and 2" across with three lobes/chambers, pointed on their distal ends; each chamber contains up to four 1/4" seeds. Both flowers and fruits are distinctive and easily recognizable.

Staphylea

Staphylea trifolia

bladdernut [Blooms: Apr-May]

Common chickweed's flower stems arise from the axils of outer pairs of leaves; the flower stems terminate in small cymes of flowers. Individual flowers are about 1/4" across, consisting of five green sepals longer than the petals, tapering to a point, hairy on the outside; five white, widely-spaced, bifid (deeply notched, each appearing double) petals; 2-10 (usually 3-5) stamens with reddish-violet anthers; and a central green ovary with three styles fused except at their flaring tips. The fruits are light brown, cylindrical, 4-6 mm long seed capsules nestled in the embrace of the (hairy) sepals with six small teeth around their upper ends. Common chickweed is usually self-pollinating (autogamous) but is occasionally cross-pollinated by flies. Chickweeds of the genus Stellaria (like common chickweed) have three styles; chickweeds of the genus Cerastium (like mouse-eared chickweed) have five styles. The flowers of whitlow-grass (Draba verna) are superficially similar to common chickweed in size and color but have only four bifid petals rather than five. One odd (but distinguishing) characteristic of common chickweed — if you bend the stem back and forth, the epidermis and parenchyma will split, leaving a central vascular bundle which will stretch elastically if you then pull the broken segments apart.

Stellaria

Stellaria media

common chickweed [Blooms: Apr-May]