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Skyblue aster's central stem terminates in a 6-18" long panicle of flowerheads; the panicle is twice as long as wide. The panicle has a central stalk (rachis), several primary branches and numerous secondary (off the primary) and tertiary (off the secondary) branches that end in flowerheads. The branches have scale-like, 3/4" long leafy bracts spaced along their lengths. Individual flowerheads are 1/2-3/4" across with 10-25 lavender or pale blue ray florets and 20-25 disk florets. The disk florets are initially yellow, turning reddish-purple with age; they seem to me to be unusually large for an aster. The phyllaries (bracts) surrounding the base of the flowerhead are small (2 mm) and tightly appressed, each with a diamond-shaped green tip and often with a spot of blue-purple at the very tip. The fruits are bullet-shaped, 1.5-2mm long dry seeds, purplish brown, with a tuft of hairs. Skyblue aster is similar to smooth blue aster (S. laeve), but that species has sessile leaves; sky blue aster has petioles on the lower and middle leaves, only the upper leaves are sessile. Skyblue aster is also similar to Short's aster (Symphyotrichum shortii), but the latter has upper leaves with a heart-shaped base rather than smoothly tapering to a sessile attachment as in skyblue aster. Generally, skyblue aster is most easily identified by the long, narrow, arrowhead-shaped lower leaves that feel like fine sandpaper on both surfaces.

Symphyotrichum

Symphyotrichum oolentangiense

skyblue aster [Blooms: Sep – Oct]

The upper stems of hairy aster give rise to 2 foot long, 1.25 foot wide panicles with a central rachis that produces spreading or ascending primary branches and smaller secondary branches that bear the flowerheads. Each flowerhead is 1/2-3/4" across with 15-35 white ray florets and 20-40 yellow disk florets that turn reddish purple or brown with age. The bracts (phyllaries) that surround the base of the flowerhead are present in 4-5 layers; they are 3 mm long and light green with a long, darker green tip. The edges of the phyllaries are often rolled under and thus may appear needle-like. The phyllaries and the bracts attached to the panicle branches both have a tiny spine at their tip. (Note: heath aster (S. ericoides) also has such spines but has smaller flowerheads, broad, blunt phyllaries with a larger spine, and the hairs on the stems are either appressed to the stem or are ascending.) The fruits are 1-1.15 mm long, tan colored, dry seeds with 3-4 longitudinal ribs and a tuft of white hairs. After blooming is completed, rosettes of 3" long, 1" wide, spoon-shaped basal leaves develop and persist through the winter into the spring, but wither away before blooming begins again.

Symphyotrichum

Symphyotrichum pilosum

hairy aster [Blooms: Aug – Nov]

The panicles of Short's aster's flowerheads are up to 18" long and 8" across; the panicles arise from the tips of the central stem and the upper lateral branches. The flowering branches are long, widely spreading or arching, and covered with small, leaf-like bracts and stiff hairs. Individual flowerheads are 3/4-1" across. Flowerheads consist of 10-20 lavender or pale blue-violet (rarely white) ray florets and 15-23 disk florets, initially yellow but becoming reddish purple over time. The phyllaries that surround the base of the flowerheads are covered with stiff, white appressed hairs, are whitish or light green with a diagnostic dark green, diamond-shaped tip, and sometimes also with a dark green medial vein. The fruits are 2-3 mm long, oblong-shaped, with a tuft of light brown hairs at one end. The indented bases of the lower and middle leaves, the slender, wingless petiole, the hairless leaves, and the phyllaries with dark green apical diamonds together are good indicators of Short's aster. Short's aster is the commonest aster in bloom in Jackson Park in early autumn; later in the autumn it is overtaken in numbers (or at least visual impact) by New England asters.

Symphyotrichum

Symphyotrichum shortii

Short's aster [Blooms: Aug – Nov]

The upper stems and branches (some arising from upper leaf axils) of white arrowleaf aster terminate in 6-18" long panicles of flowerheads; the panicles are 2-3 times longer than wide. Each flowerhead is 1/2-3/4" across with 8-15 pistillate, white (rarely pale lavender) ray florets and 10-15 perfect (both stamens and styles present) disk florets. The disc florets are initially yellow but turn pink or light purple with age such that the flowerheads with yellow disk florets are often in the minority on the plant. The individual disk florets strike me as being larger than on many asters. The phyllaries (bracts) covering the base of the flowerhead are narrowly lance-shaped, usually hairless, dark green in the middle tapering to narrow, stiff, green to purplish tips. The fruits are 2 mm long, purplish-brown dry seeds with longitudinal ribs and a tuft of hairs at one end. White arrowleaf aster is similar to Drummond's aster (Symphyotrichum drummondii) but is less hairy and has distinctly different phyllaries. Among asters with white flowerheads, only white arrowleaf aster has heart-shaped leaves. (Other asters with heart-shaped leaves typically have blue to violet flowers.)

Symphyotrichum

Symphyotrichum urophyllum

white arrowleaf aster [Blooms: Aug/Sep – ?]

Some upper stems produce compound umbels at their tips made of tiny, yellow flowers; the umbels are very open (they remind me of a fireworks burst), up to 4-7" across with 12-15 umbellets of about a dozen flowers. Individual flowers are about 1/8" across with five tightly furled, yellow petals; five stamens emerging from between the incurled petals and widely spreading with pale yellow anthers; and two styles. Sepals are absent. The flowers tend to be either pistillate (stamens reduced or absent) or staminate (styles reduced or absent), but a few perfect flowers occur. Yellow pimpernel prefers dry habitats on a slope at the edge of woods.

Taenidia

Taenidia integerrima

Yellow pimpernel [Blooms: May-?]

Purple meadow-rue is a native perennial that is listed as "vulnerable" in Illinois. The species is found in most of the lower 48 states in the U.S. but is notably absent from states on the Atlantic and Pacific coasts (however, it is present along the Gulf of Mexico). Purple meadow-rue is dioecious — some plants produce only staminate (male) flowers, each with around a dozen stamens; other plants bear only pistillate (female) flowers, each with about ten pistils. The male flowers are the showier of the two. Petals are absent (the plant is wind-pollinated). Both the narrow, whitish-grey, thread-like pistils and the yellow-green stamens (with pale yellow anthers) droop from 4-5 light green sepals (that may drop off). Fertilized female flowers develop a single, spindle-shaped fruit that is 1/4" long, elongated, sharply pointed with a remnant style, 4-6 raised ribs, and slightly concave along one side. The fruit is initially green, turning brown with maturity. Early meadow-rue (Thalictrum dioicum) is similar to purple meadow-rue but has light purplish-green stems and leaves with 3-9 (typically five) rounded lobes and long petioles on the upper leaves.

Thalictrum

Thalictrum dasycarpum

purple meadow-rue [Blooms: May-Jun]

Early meadow-rue's main stem terminates in a panicle up to a foot long and wide; each branch in the panicle has a corymb of about five flowers at its tip. Early meadow-rue is dioecious — some plants produce only staminate (male) flowers, each with 10+ stamens; other plants bear only pistillate (female) flowers, each with up to 15 pistils. Both male and female flowers droop from their stems, but the male flowers have a longer droop than do the female flowers. Both male and female flowers are about 1/2" long and 1/4" across with 4-5 pale green sepals with white margins (which often fall off). Neither male nor female flowers have petals; pollination is via the wind. Fertilized female flowers produce ellipsoid, dry seeds that are pointed at both ends; the surface bears strong ribbing. Early meadow-rue prefers shady woodlands. It has more lobes on its leaves (usually 3-5), and the lobes are more rounded than purple meadow-rue (Thalictrum dasycarpum), the other Thalictrum species in Jackson Park, does (2-3 pointed lobes).

Thalictrum

Thalictrum dioicum

early meadow-rue [Blooms: Apr-May]

Field pennycress's central stem and the upper side stems produce racemes of flowers at their tips. The flowers bloom at the tip of the raceme, which continues to elongate to 3-8" long and to produce new buds; the prominent, disk-shaped fruits develop lower on the raceme. Individual flowers are about 1/8-1/4" across on a 1/2" pedicel; there are four green sepals with whitish edges, four white, oval petals, six yellowish-green stamens (three on each side of the flattened ovary), and a single short style. The fruit is a 1/2" wide flattened, winged disc with a distinct U-shaped notch at the apex; the fruits are borne on the flower pedicels, which spread perpendicular to the stem. Each seedpod has two compartments, each compartment containing several dark brown seeds. Other mustards have long, cylindrical seedpods; the nearly-circular seedpod of field pennycress is unique among Jackson Park mustards. Young field pennycress plants are edible but become very bitter at flowering. Cows that feed on field pennycress produce "tainted" milk that tastes like garlic.

Thlaspi

Thlaspi arvense

field pennycress [Blooms: Apr-Jun]

Erect hedge-parsley's main stem and side stems give rise to umbels 1.5-2" across of 10-20 umbellets of flowers. Each flower is about 1/8" across consisting of five white petals (often markedly unequal in size), and a creamy white center surrounded by five pink to white stamens. The fruit is an 1/8" long oval covered in hooked hairs to aid in bur-like dispersal; it contains two seeds. (The fruit of common hedge-parsley (Torilis arvensis) has straight or only slightly curved hairs, not hooked, but the hairs are barbed.)

Torilis

Torilis japonica

erect hedge-parsley [Blooms: Jun-Jul]

Ohio spiderwort's flowers and buds occur in a cluster, each cluster on a hairless flower stalk (pedicel) with a pair of long, leaf-like bracts at its base at the apex of the plant. Individual flowers are about 1-1.5" across, with three green, oval sepals offset from the petals; three rounded blue-violet petals; six stamens with blue filaments and bright yellow anthers; a single blue style with a small white stigma; and fine, thread-like, violet hairs radiating out from the stamen and style bases in the center of the flower. The seed capsules are 4-8 mm long, ovoid, with three distinct compartments. The mature seed capsule splits into three sections, each releasing 3-6 oblong, strongly ribbed, brown seeds. The flowers open early in the morning but close in the afternoon (later on cloudy days). Ohio spiderwort is the commonest spiderwort in Illinois and the only Tradescantia species to occur in Jackson Park; the very similar Virginia spiderwort (Tradescantia virginiana) can be distinguished from Ohio spiderwort by its hairy flower stalks (pedicels).

Tradescantia

Tradescantia ohiensis

Ohio spiderwort [Blooms: Apr-Sep]

Western goat's-beard's stems terminate in a single flowerhead, with the stem strongly swollen beneath the flowerhead. Individual flowerheads are pale lemon yellow and about 2" across. Between 50-180 ray florets are present; disc florets are absent. A series of about 13 sharply pointed, narrow bluish-green bracts (significantly longer than the outer ray florets) surround the base of the flowerhead. The outer ray florets (the "petals") are much longer than the inner ray florets. All ray florets have an outer tip of the "petal" decorated with five small teeth. Near the base of the ray florets is a composite reproductive structure consisting of a single style, bifurcated at its tip, and five stamens with black anthers closely appressed to the middle of the style. The flowerheads open in the morning but close in the afternoon; the flowerheads sometimes track the sun like a sunflower. Fertilized florets produce a long, narrow, naked seed densely covered with small teeth; the seed is attached to a plume of fine white hairs (like a dandelion). The seed head looks like a giant version of a dandelion seed head. Western goat's beard is very similar to yellow goat's beard (Tragopogon pratensis) but prefers drier habitats; the latter has only eight bracts (rather than 13) surrounding the flowerhead and the bracts are the same length (not longer) than the outer ray florets.

Tragopogon

Tragopogon dubius

western goat's-beard [Blooms: May-Sep]

Yellow goat's-beard stems produce a long naked stalk from their tips that bears a single flowerhead. Individual flowerheads are bright yellow and 1-2.5" across. Between 50-180 ray florets are present; disc florets are absent. A series of about eight sharply pointed, narrow green bracts (the same length or shorter than the outer ray florets) surround the base of the flowerhead. The outer ray florets (the "petals") are much longer than the inner ray florets. All ray florets have an outer tip decorated with five small teeth. Near the base of the ray florets is a composite reproductive structure consisting of a single yellow style, bifurcated at its tip, and five stamens with black anthers closely appressed to the middle of the style. Fertilized florets produce a long, narrow naked seed; the seed is attached to a plume of fine white hairs (like a dandelion). The seed head looks like a giant (3" across) version of a dandelion seed head. Like Tragopogon dubius, it's an exotic, but at least it's less aggressive. Distinguish yellow goat's beard (T. pratensis) from western goat's beard (T. dubius) by flower color (bright yellow vs. pale yellow), foliage color (green vs. bluish green, number of floral bracts (8 vs. 13), leaf shape (tips curled vs. tips straight), and length of the floral bracts (same as petals vs. much longer than petals).

Tragopogon

Tragopogon pratensis

yellow goat's-beard [Blooms: May-Aug]

Each side stem of yellow bellwort produces a single, elongate bell-shaped flower at its tip. The flowers droop on their stem-like pedicels (about 1 " long) such that they open downwards and are often partially hidden by the leaves. Each flower is 1-2" long and consists of six yellow tepals twisted along their length (often with greenish bases), six white stamens with elongated yellow anthers, and a pistil with a three-lobed stigma. The fruit is a three-section capsule that looks (vaguely) like three fat triangles with oval bases, stuck base to base to base. Seeds are freed when the capsule splits. Seed dispersal is added by the attraction of ants to a fleshy, nutrient-rich appendage (an elaiosome) on the seed; the ants carry the seed back to their nest where they eat the elaiosome and abandon the seed.

Uvularia

Uvularia grandiflora

yellow bellwort [Blooms: Apr-May]

The apex of the common mullein plant is a club-shaped flower spike 6-24" long, covered with scattered light-yellow flowers each 3/4" across. Individual flowers have five hairy, green sepals; five pale yellow petals (the lower two slightly larger than the upper three); five stamens with reddish-orange anthers; and a single green style with a heart- or disc-shaped stigma. The upper three stamens are shorter than the other two stamens and covered with white or yellow hairs; the lower two stamens are longer and largely hairless. The fruit is an oval, two-celled seed capsule, brown when mature, each cell filled with numerous (25-150) tiny orange to brown seeds. Common mullein is immediately recognizable and impossible to confuse with any other local plant.

Verbascum

Verbascum thapsus

common mullein [Blooms: Jun-Aug]

The top of the wingstem plant is a dome (a panicle) of coarse-looking flowerheads. Each flowerhead is 1-2" across, with 2-10 yellow, oblong, drooping petals, 1/2-1" long that are derived from the sterile ray florets. Some 40-60+ yellowish-green, tubular disk florets project in all directions in a hemisphere of widely-spaced, robust, bud-like blunt cylinders, the tips of the disk florets. When receptive to pollen, the disk florets bear a protruding style with a bifurcated, curling pair of stigmas (likened by one source to a "pince-nez" — a pair of eyeglasses on a handle); the five stamens and their flattened, black anthers are appressed to the body of the style. The fruit is a teardrop-shaped seed with a pair of thin wings and two short, robust, spiky awns. Immature seeds are green; the seeds (and seedheads) turn brown when mature.

Verbesina

Verbesina alternifolia

wingstem [Blooms: Jul-Oct]

Riverbank grape's flowers develop in 5" long, drooping, cylindrical or pyramidal panicles opposite the leaves of new branches (but skipping every third leaf). The flowers are greenish-yellow and are best described as skeletal; they may be either unisexual or (rarely) perfect (both male and female reproductive organs functional). The male and female unisexual flowers usually both occur on the same plant, either in separate or mixed clusters; both male and female flowers are 1/8" across or less, with a minute calyx and five green to yellowish petals that quickly fall off. Male flowers have five very long, pale, erect stamens around a button-like center; female flowers have an ovary with a short style and five short, sterile stamens, often contorted. The fruit are .... grapes — a round berry, 1/4-1/2" across containing 1-4 seeds, initially green but turning blue-black as it matures, often with a white bloom on the skin. You know, grapes. The fruit of riverside grape is often sour until after a frost, then becomes more "sweet/tart" to the tongue. Riverside grape is subject to parasitism by the grape filbert gall midge (Ampelomyia vitiscoryloides), an insect whose larvae produce large, very hairy, conical galls at leaf nodes. This gall midge is restricted to the eastern half of North America; it was first described by Alpheus Spring Packard, Jr. in 1869.

Vitis

Vitis riparia

riverbank grape [Blooms: [May-Jun]

Common cocklebur produces a spike-like raceme of compound flowerheads, shorter than the leaf petioles, that arises from the axils of each upper leaf and from the tip of the central stem. Each raceme produces exclusively male flowerheads in its upper parts and exclusively female flowerheads in the lower regions. The male flowerheads are about 1/4-3/4" across, elevated on a short pedicel on top of 1-3 series of white floral bracts. Each of the approximately 20 tiny, brownish-green staminate florets (nominally disk florets) consist of a tubular 1-2 mm long corolla; a minute, non-functional ovary; and five stamens with fused filaments (like a tree trunk) but with their white anthers free, positioned in a ring like a little palm tree or, possibly, a hand. The staminate florets quickly shed their pollen and drop off. The female flowerheads (also nominally disk florets) are 3/4-1.5" long and two-thirds as wide, stalkless, each (usually) containing two female florets nearly enclosed by floral bracts covered with curly white hairs that have a bur-like appearance from the dense, hooked prickles covering their surface. At the tip of the flowerhead are two spines, longer and much stouter than the prickles, each with a hole from which protrudes the divided style of a female floret. There is no corolla in the female florets. The florets are wind pollinated, and their styles quickly wither away after fertilization. Each female floret produces a single, oblong, dark gray to black seed 3/8-1/2" long, tapering to a point at each end and covered by dark membranes, all buried in the bur-like bract. The female flowerhead stiffens through the autumn, turns brown, and overwinters. The seeds are highly poisonous, containing toxic diterpene glycosides. One of the contained seeds germinates the following year; the other delays germination for two years. Common cocklebur prefers wet, sandy soils.

Xanthium

Xanthium strumarium

common cocklebur [Blooms: Sep-?]

The tips of the upper stems of heartleaf alexanders produce 2-3" wide compound umbels of 7-15 umbellets, each with 10-20 tiny yellow flowers. The central flower of each umbellet is sessile; the rest have short pedicels (a diagnostic character of the genus). Each 1/8" wide flower consists of five yellow petals that remain curled, five stamens with yellow anthers projecting through the spaces between the petals, and two styles. The fruits are a ribbed capsule with a flat top from which the remnants of the styles project. Heartleaf alexanders is restricted to northeastern Illinois, where it is uncommon; it is usually found in drier habitats than its much more common relative, golden alexanders (Zizia aurea).

Zizia

Zizia aptera

heartleaf alexanders [Blooms: Apr/May–?]