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Black-eyed susan flowerheads are 2-3" across; they have two series (outer longer than inner) of numerous slender, sharply-pointed, green bracts beneath the flowerhead, each densely covered with long, stiff hairs; 8-20 (usually less than 12) yellow-orange, sterile ray florets (rarely with a maroon patch near the base of the petal); and 250 to over 500 dark purplish-brown disk florets, each with slender, sharply pointed styles. The fruit consists of numerous four-sided brown or black seeds that remain embedded in the remnants of the disc flowers (a gumdrop shaped, brown mass).

Rudbeckia

Rudbeckia hirta

black-eyed susan [Blooms: Jun-Oct]

Cutleaf coneflower (aka, green-headed coneflower) is an imposing tallgrass prairie plant, 3-8' tall, that can form dense stands. The composite flowerheads are 2-3" across. Eight to fifteen light green, oblong bracts lie beneath a central globe-like, light green to yellow cone surrounded by 6-12 yellow, drooping, sterile ray florets. The disk florets have yellow to yellowish green corollas and a protruding bifurcated stigma; the stamens are yellow, fused to the base of the corolla, with deep purple anthers attached around the styles side-by-side. Immature disk florets are green (thus the alternative common name); mature disk florets produce a spikey appearance. The disk florets mature from the bottom of the cone to the top, lending a pincushion-like appearance to the cone.

Rudbeckia

Rudbeckia laciniata

cutleaf coneflower [Blooms: Jul-Sep]

Sweet coneflower flowerheads are 2-3.5" across, occurring as single blooms on the tips of stems or branches. A flowerhead is comprised of numerous 1/2" long medium green, lance-shaped phyllaries, downy with short hairs; 10-20 bright yellow, oblong, sterile ray florets with a tiny notch at their tips; and a deep brown hemisphere of 200-400 disk florets (the central disc). The disk florets have a tubular, dark reddish-purple corolla; bifurcated stigmas that have sharp tips; with the purplish-brown anthers adhering side by side around the deep purple styles. Sweet coneflower is distinctive among the Jackson Park coneflowers with its chocolate brown disk and bright yellow, elongate-oval petals with a shallow notch at their tips.

Rudbeckia

Rudbeckia subtomentosa

sweet coneflower [Blooms: Jul-Aug]

Brown-eyed susan is a native bushy plant that branches at the leaf axils and produces prodigious numbers of small flowers, up to 5' tall. The stems are deep reddish brown and covered with long white hairs. The leaves are alternate, twice as long as wide, lance-shaped, up to 4" long and covered by short, stiff hairs that give it a rough texture. Some lower leaves are three-lobed, sometimes deeply, and halberd- or trident-shaped. Upper stems terminate in one or two flowerheads, 1-2" across. Each flowerhead has an underlying inner and outer series of eight (total) green bracts with hairs along the margins; 6-13 golden-yellow, sterile ray florets, often with a small notch in the tips of the oval petals; and a central cone of 150 to over 300 purplish-brown, cylindrical disc florets with five stamens and short, blunt, bifurcated stigma. If in doubt, brown-eyed susan tends to be taller than black-eyed susan, with fewer petals (6-13 vs. 8-20) and smaller flowers.

Rudbeckia

Rudbeckia triloba

brown-eyed susan [Blooms: Jun-Sep]

Pale dock produces an inflorescence 4-12" long at the tip of the stem; branches in the inflorescence are few but are densely covered with numerous whorls of 10-20 small (1/8-1/4" long), green or yellow-green flowers drooping on 1/8-1/4" long pedicels. Individual flowers have six green tepals (three sepals and three similar petals) that spread at flowering to expose either six stamens or three styles, but close again after pollination. (Flowers are either pistillate (female) or staminate (male) and are wind pollinated.) After fertilization of the pistillate flowers, the fruit — a single seed in the middle of three equally-spaced wings — replaces the flower. Pale dock's seeds can persist in the soil for at least ten years.

Rumex

Rumex altissimus

pale dock [Blooms: May-Jun]

The inflorescence at the tip pf the stem in curly dock produces branching clusters with numerous whorls of 10-25 flowers dangling from slender pedicels. Flowers are tiny (1/8-1/4"), red- or yellow-green, with six tepals (three sepals and three very similar petals), three styles, or six stamens, or both; the flowers droop from a 1/4" pedicel. (The flowers are either pistillate (female), staminate (male), or perfect (both male and female organs). The flowers are wind pollinated. The tepals spread out when the flowers are mature, exposing the stamens and/or styles, then close up after pollination. The fruit is a single seed sandwiched in the center of three heart-shaped, papery membranes (derived from the tepals) set at 120° from each other, dark brown when mature. Note that more mature curly dock fruit can take on a distinct reddish tint, superficially looking like bitter dock fruit, but curly dock fruits will never show the long tooth- or finger-like projections from the edges of the wings that bitter dock exhibits.

Rumex

Rumex crispus

curly dock [Blooms: May-Jun]

The inflorescence of bitter dock is a panicle of whorled racemes up to 1' long. The greenish-red flowers droop from 1/2" pedicels. Individual flowers are 1/4" long with six tepals (six sepals and six similar petals) with pistillate (female) and male (staminate) flowers mixed in each inflorescence. Pistillate flowers have a single pistil, three styles with fringe-like, divided stigmas, and inner tepals that are (often) red; staminate flowers have six stamens and dull yellow inner tepals. After fertilization, the inner tepals of the pistillate flowers enlarge and surround an expanding, hard-shelled, three-sided seed with three membranous wings, each wing with 2-4 spiny teeth along their free edges. The fruits turn bright red as they mature.

Rumex

Rumex obtusifolius

bitter dock [Blooms: May-Jul]

Common arrowhead's flowering stalks are about as tall as the leaves, either angular or round in section. Branches of the inflorescence end in 2-5 whorls of 2-3 flowers; the terminal branch bears 3-9 whorls of 2-3 flowers. Beneath each whorl 2-3, 1/8-1/3" long, boat-shaped, green floral bracts unite. Male (staminate) flowers have 25-30 yellow stamens while female (pistillate) flowers have a central green sphere of carpels; the male flowers are located above the female flowers in each raceme. Perfect flowers (both male and female organs present) do occur but are rare. Individual flowers are about 1-1.5" across with three oval, green sepals and three rounded, white petals about twice the length of the sepals. Male flowers have 7-30 yellow stamens in the center of the flower; female flowers instead have a bur-like green cluster of hundreds of carpels and their spike-like pistils (which have terminal or lateral styles). The pistils form a dense spiral on the surface of the central green mass. After fertilization, the female flowers lose their petals and transform into a bur-like, brown fruit about 3/4" across when mature; the spikes on the bur are individual seeds (achenes) that have a large, fang-like projection at right angles to the main axis of the seed.

Sagittaria

Sagittaria latifolia

common arrowhead [Blooms: Aug – Sep]

Upper branches of elderberry end in flat to domed panicles of flowers 3-10" across. Individual flowers are about 1/4" across, with a green, cup-shaped calyx, a creamy white corolla with five petal-like lobes, five long, spreading stamens with white anthers, and a creamy white hemispherical ovary with 3-5 short styles; flowers occur in umbel-like panicles 10" across made of dozens of flowers. The fruits are <1/4" wide spherical drupes in starburst-like sprays, dark purple to black when mature. The flower stalks are light green, turning purplish as the fruit ripens. The raw fruit is sweet with a slightly bitter or astringent aftertaste but may induce nausea or diarrhea if eaten in more than small quantities. The fruit is widely used in a variety of herbal supplements (as infusions and extracts), jam, and herbal wine. Note that all parts of the plant including the sap (except the flesh of the fruit) contain toxic cyanide-producing glycosides; berries should be cooked before consumption.

Sambucus

Sambucus nigra

elderberry [Blooms: May-Jul]

Bloodroot flowers are 1.5-3" across with 8-16 (typically 8) bright white petals, numerous stamens with golden yellow anthers, and an oval, green pistil with a pale yellow, two-lobed stigma at its tip. Two light green sepals are present in the floral bud but they drop off as soon as the flower begins to open. After fertilization, the ovary expands into a spindle-shaped seed capsule about 1" long, yellow when mature, that splits to release the seeds. The seeds are disbursed by ants that are attracted by the white, fleshy appendage (an elaiosome) on the bright red seed. The red sap is highly toxic, both externally and internally — it inhibits the cellular Na+/K+ ATPase enzyme and kills the cells. Avoid getting the sap on your skin and certainly avoid getting it in your eyes or on mucous membranes.

Sanguinaria

Sanguinaria canadensis

bloodroot [Blooms: Mar-Apr]

The tips of the upper stems of Canadian black snakeroot give rise to umbels 2.5" across of greenish white flowers; each umbel usually consists of 1-4 umbellets, each with 2-3 pistillate flowers or perfect flowers and 2-6 staminate flowers. In perfect flowers, the pistils develop (and are deployed) before the stamens develop; the pistillate stage persists for much longer. Each flower has a green calyx with five lance-like lobes slightly longer than the five greenish-white petals. Male and perfect flowers add five stamens with white anthers and lack the series of hooked spines of the pistillate flowers. Female and perfect flowers have ovaries densely covered with distinctive, neatly stacked series of long, hooked bristles arrayed perpendicular to the axis of the flower. The fruit is round, 3-5 mm long, covered in rows of those same hooked bristles; it splits on maturity to release the two seeds.

Sanicula

Sanicula canadensis

Canadian black snakeroot [Blooms: Jun-Jul]

On black snakeroot, one to five 1/2" wide globoid umbels (flower clusters) arise from the tips of the upper stems. Each umbel is packed with 20-60 tiny flowers and their flower stalks (pedicels). In each umbel, 1-3 flowers are "perfect" (with both male and female organs) and the rest are staminate (stamens only). In addition to the male stamens, the perfect flowers have two long, cylindrical styles (emerging from a hidden ovary) that arch and recurve from the middle of the flower; the styles are much longer than the hooked bristles on the ovary. Both staminate and perfect flowers have a minute (<1 mm) green calyx with five rounded teeth, five dull yellow to greenish-yellow, heart-shaped petals (tip of the heart towards the center of the flower), and five protruding stamens <3 mm long with obvious yellow (or brown) anthers. The fruit is a 1/4" long round to cylindrical bur covered with short, hooked bristles; the remains of the styles often persist. The bur splits to release the two seeds inside.

Sanicula

Sanicula odorata

black snakeroot [Blooms: May-Jun]

Alpine squill is a 4-8" tall early spring plant, native to Eurasia. Alpine squill has two (rarely three or four) 3-6" long, narrow, curved, fleshy leaves. One to six red-brown, hairless, leafless flower stalks arise from between the basal leaves; each stalk supports 2-10 deep blue, erect flowers less than 1/2" across on individual, hairless pedicels. The basal leaves clasp the flower stalks up to about half their length. Each flower faces upwards (unlike Scilla siberica), has six spreading tepals, six stamens with dark blue anthers, and a single pistil. The stamens are distinctive in that the filaments are flattened along the flower's radial axis and taper distally; from the side they look like isosceles triangles with the anthers on their tips. Alpine squill contains cardiac glycosides; all parts of the plant are highly toxic.

Scilla

Scilla bifolia

alpine squill [Blooms: Mar-Apr]

Siberian squill is another of the early spring ephemerals native to the Caucasus and west Asia that has been wildly successful in northern Illinois and parts of Minnesota, displacing native species. Its leaves are restricted to a basal set of 2-3 leaves about 5" long and 1/4-1/2" wide, linear with a blunt tip, a shiny green surface, and smooth margins. The leaves tend to roll inward along their margins with the curl increasing towards the tip of the leaf. One or more purplish-green, 6" tall flower stalks arise from the center of the ring of basal leaves with single (rarely two or three) blue, nodding flowers on each stalk, suspended from a slender pedicel that arises directly from the unbranched flower stalks. Each flower is about 1" across with six flaring blue (rarely, white) tepals (each typically with a dark blue line down the middle), six white stamens with dark blue anthers, and a single green pistil with an undivided blue style. Siberian squill can be confused with alpine squill (Scilla bifolia), but in alpine squill the flowers face laterally or upwards rather than drooping like Siberian squill, and the flower stalks are distinctly red-brown (rather than brownish green) and branched rather than supporting flowers with a simple pedicel. Despite the common name, Siberian squill is not native to Siberia; it is native to a broad swath of western Europe from Turkey through the Caucasus and southwestern Russia. Sources differ on whether Siberian squill is toxic but note that it is avoided by vertebrate herbivores; proceed with caution.

Scilla

Scilla siberica

Siberian squill [Blooms: Mar-Apr]

Biting stonecrop (aka, mossy stonecrop) is an exotic succulent that is native to Greenland and western Europe but naturalized across most of the U.S. and the lower Canadian provinces. It is a mat-forming plant with creeping stems that roots at nodes; the stems produce short, erect vegetative and flowering stems. The leaves are alternate, sessile, and densely imbricated on the erect stems; they are green or yellowish-green, with old, white leaves persisting. The succulent-like leaves are 4-5 mm long, 2.5-3 mm across, and 1.5 mm thick, convex on both the upper and underside and blunt-tipped. The inflorescence arises from the tip of one of the erect stems; it is about 5/8" (1.5 cm) long, 1/4-1.25" (5-30 mm) across, with 1-3 branches, each with 1-4 sessile (or on 1 mm long pedicels) flowers with a basal bract. The sepals closely resemble the leaves. The flowers have five bright yellow petals, typically 10 stamens with yellow anthers, and 4-5 yellow pistils, each with a single style and a pinhead-shaped stigma. The fruit is a five-pointed yellowish star of united, inflated follicles filled with numerous seeds. Biting stonecrop is not a good competitor; it grows best in thin, dry soils like cliffs, shingle, and beaches. However, it is tough — it stores water in its leaves and reportedly can go six months without external water.

Sedum

Sedum acre

biting stonecrop [Blooms: Jun]

The tips of the upper stems of common groundsel produce dense clusters of flowerheads on short peduncles; inflorescences can also arise from leaf axils. Common groundsel flowerheads are cylindrical, 8 mm long and 4 mm across, with black-tipped, green bracts around the outside at both the base and the apex and 13-80 yellow disc florets peeking out the end; mature flowerheads are easily mistaken for flower buds. The disc florets have tubular yellow corollas, five stamens fused to the upper part of the corolla with yellow-brown anthers, and a bifurcated style with purplish stigmas. There are no ray florets. This is an easy plant to overlook. The flowerheads are twice as long as wide. The seedhead looks like a compact dandelion tuft with seeds on parachutes of light brown hairs.

Senecio

Senecio vulgaris

common groundsel [Blooms: May-Jul]

The five petals and five sepals of Maryland senna's 3/4" flowers are both yellow. The single pistil has many spreading white hairs and arches upward. The anthers of the 10 stamens are a striking dark brown (like partridge pea). The upper three stamens have short filaments and short anthers that produce only sterile pollen (as a pollinator reward). The middle four stamens have short filaments and long anthers while the lower three stamens have long filaments and long anthers; all seven of these stamens produce fertile pollen that is released by "buzz pollination." The flowers do not produce nectar. The fruit is a flattened seedpod 1/2" across and 3-4" long with 10-18 segments, each containing a single 1/4" flattened seed.

Senna

Senna marilandica

Maryland senna [Blooms: Jul]

Flower buds arise singly or in small groups on 1/2" long peduncles from the leaf axils. Because the flower peduncles are shorter (1/2") than the leaf petioles (1"), the flowers are often partly hidden among the leaves. Individual flowers are about 8 mm across. A flower contains five green, triangular sepals; five light yellow or light orange, rounded, floppy petals with pale lines that function as nectar guides and asymmetrical tips; and a central reproductive column (like mallows, to which prickly sida is related). That column consists of five styles fused along most of their lengths, each with a globose terminal stigma, and numerous stamens with yellow anthers. After fertilization, a circular seedpod forms from the five carpels; when mature, the seedpod turns brown and breaks into five segments, each containing a single reddish-brown, three-angled seed.

Sida

Sida spinosa

prickly sida [Blooms: Aug-Sep]