Senna marilandica
Maryland senna [Blooms: Jul]
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The five petals and five sepals of Maryland senna’s 3/4″ flowers are both yellow. The single pistil has many spreading white hairs and arches upward. The anthers of the 10 stamens are a striking dark brown (like partridge pea). The upper three stamens have short filaments and short anthers that produce only sterile pollen (as a pollinator reward). The middle four stamens have short filaments and long anthers while the lower three stamens have long filaments and long anthers; all seven of these stamens produce fertile pollen that is released by “buzz pollination.” The flowers do not produce nectar. The fruit is a flattened seedpod 1/2″ across and 3-4″ long with 10-18 segments, each containing a single 1/4″ flattened seed.
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Maryland senna is almost identical to wild senna (Senna herbecarpa) but has a less exuberantly hairy pistil than wild senna. I have seen Maryland senna in Jackson Park over several years, but I have yet to find wild senna. Maryland senna is a sparingly branched but still bushy plant up to six feet tall. The stout stem is green, round in section, and either ribbed or angular (but with rounded corners). The leaves are alternate, even-pinnate, with 6-12 pairs of bluish-green leaflets 1/2-1″ across and 1-2.5″ long with pointed tips and a basal 1/8″ long petiolule that attaches to the rachis of the compound leaf. The leaf petiole is 2-6″ long, grooved along the upper surface. Near the base of the leaf petiole, on its upper side, is a budlike gland (grayish-purple and dome-shaped) that functions as an extrafloral nectary to attract ants and ladybird beetles (presumably for protection, rather like the extrafloral nectaries of partridge pea, Chamaecrista fasciculata). The base of the petiole is swollen; the swollen structure (a pulvinus) permits movement and reorientation of the leaf.
More information
Maryland senna is almost identical to wild senna (Senna herbecarpa) but has a less exuberantly hairy pistil than wild senna. I have seen Maryland senna in Jackson Park over several years, but I have yet to find wild senna. Maryland senna is a sparingly branched but still bushy plant up to six feet tall. The stout stem is green, round in section, and either ribbed or angular (but with rounded corners). The leaves are alternate, even-pinnate, with 6-12 pairs of bluish-green leaflets 1/2-1″ across and 1-2.5″ long with pointed tips and a basal 1/8″ long petiolule that attaches to the rachis of the compound leaf. The leaf petiole is 2-6″ long, grooved along the upper surface. Near the base of the leaf petiole, on its upper side, is a budlike gland (grayish-purple and dome-shaped) that functions as an extrafloral nectary to attract ants and ladybird beetles (presumably for protection from herbivores, rather like the extrafloral nectaries of partridge pea, Chamaecrista fasciculata). The base of the petiole is swollen; the swollen structure (a pulvinus) permits movement and reorientation of the leaf. The five petals and five sepals of the 3/4″ flowers are both yellow. The single pistil has many spreading white hairs and arches upward. The anthers of the 10 stamens are a striking dark brown (like partridge pea). The upper three stamens have short filaments and short anthers that produce only sterile pollen (as a pollinator reward). The middle four stamens have short filaments and long anthers while the lower three stamens have long filaments and long anthers; all seven of these stamens produce fertile pollen that is released by “buzz pollination.” The flowers do not produce nectar. The fruit is a flattened seedpod 1/2″ across and 3-4″ long with 10-18 segments, each containing a single 1/4″ flattened seed. Maryland senna is superficially similar to partridge pea: (1) both have even-pinnate compound leaves, but partridge pea has 5-18 pairs of small (~2/3″ long) leaflets while Maryland senna usually has 6-12 pairs of larger (1-2″ long) leaflets, and (2) both have yellow flowers with dark brown anthers, but the flowers of partridge pea are markedly asymmetrical with one petal twice the size of the other four and another petal that curls around the stamens, while the flowers of Maryland senna are bilaterally symmetrical (one side is the mirror image of the other).
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![Evening primrose — clusters of yellow flowers occur at the apex of the plant. The flowers are 1-2" across, with four 1/2-1" long green sepals strongly reflected over a long (3/4-2"), green, cylindrical floral tube (the ovary is located just above the leaf axil), four heart-shaped yellow petals, eight stamens, and a single style with an X-shaped stigma. The sepals typically remain connected (in pairs) along one edge until the flower opens. The seedpods develop within the ovary (the base of the elongate floral tube); they are hairy, urn-shaped with rounded edges, and have four or eight small lobes on the apex. The seeds are claimed to remain viable in the soil for up to 70 years. The flowers open at dusk and close early in the day. — — — There are (at least) three evening primroses in Jackson Park — common evening primrose (Oenothera biennis), northern evening primrose (Oenothera parviflora), and hairy evening primrose (Oenothera villosa). They are easily confused with each other. (Judging by the conflicts between the accounts on Illinois Wildflowers [https://www.illinoiswildflowers.info/], Minnesota Wildflowers [https://www.minnesotawildflowers.info/], and Missouri Plants [http://www.missouriplants.com/], even the experts are confused.) All three species have yellow flowers with an X-shaped stigma, reflected sepals, petals with a small notch in their apex, stems that may be red or green, and lance-shaped leaves with short, widely-spaced teeth. Northern evening primrose is distinguished by (1) the presence of a knob or ridge at the tip of each sepal which the other two species lack (2) a seed capsule with four robust, flaring lobes with rounded tips at the apex of the seed capsule, and (3) a stigma that is very robust and almost fills the opening of the flower. Northern evening primrose has green to yellowish-green sepals. Hairy evening primrose (often but not always) has (1) striped or reddish-tinged sepals (check the buds where this character is often easier to see), (2) flowers that turn orange as they begin to senesce (although the reliability of this character is uncertain), and (3) always has glandular hairs covering the ovary that have "pimple-like" bright-red bases; these red pimples may also occur on the stems. All three species are somewhat similar to prairie sundrop (O. pilosella); the latter has very hairy stems and leaves, and translucent lines that radiate from the base of the petals in the flowers.](https://jacksonparkwildflowers.info/wp-content/uploads/2025/03/Oenothera_biennis-1t-300x300.jpg)
![Northern evening primrose flowers are 3/4-1.5" across, have four yellow, heart-shaped petals, eight yellow stamens, and a cross-shaped stigma on a modest style; the stamens are as long or longer than the petals and the stigma is nearly fills the floral opening. At the base of the flower are four sepals 1/2-1" long, that bend back along the 3/4-1.5" long floral tube as the flower develops with pairs of sepals attached along one edge until the flower opens. Only northern evening primrose with a distinct ridge or knob near the tips of the sepals. The seed capsule develops from the ovary; it is 3/4-1.5" long, cylindrical but tapering slightly towards the tip, with four robust, flaring lobes at the tip. — — — There are (at least) three evening primroses in Jackson Park — common evening primrose (Oenothera biennis), northern evening primrose (Oenothera parviflora), and hairy evening primrose (Oenothera villosa). They are easily confused with each other. (Judging by the conflicts between the accounts on Illinois Wildflowers [https://www.illinoiswildflowers.info/], Minnesota Wildflowers [https://www.minnesotawildflowers.info/], and Missouri Plants [http://www.missouriplants.com/], even the experts are confused.) All three species have yellow flowers with an X-shaped stigma, reflected sepals, petals with a small notch in their apex, stems that may be red or green, and lance-shaped leaves with short, widely-spaced teeth. Northern evening primrose is distinguished by (1) the presence of a knob or ridge at the tip of each sepal which the other two species lack (2) a seed capsule with four robust, flaring lobes with rounded tips at the apex of the seed capsule, and (3) a stigma that is very robust and almost fills the opening of the flower. Northern evening primrose has green to yellowish-green sepals and is the only one of the three evening primroses to (sometimes) have reddish color on some or all of the midvein of the leaves. Hairy evening primrose (often but not always) has (1) striped or reddish-tinged sepals (check the buds where this character is often easier to see), (2) flowers that turn orange as they begin to senesce (although the reliability of this character is uncertain), and (3) always has glandular hairs covering the ovary that have "pimple-like" bright-red bases; these red pimples may also occur on the stems. All three species are somewhat similar to prairie sundrop (O. pilosella); the latter has very hairy stems and leaves, and translucent lines that radiate from the base of the petals in the flowers.](https://jacksonparkwildflowers.info/wp-content/uploads/2025/03/Oenothera_parviflora-1t-300x300.jpg)
![Hairy evening primrose produves an elongating spike of flowers arises from the apex of the stem; flowers bloom near the tip while older flowers and seed capsules develop below. Individual flowers are 3/4-1.5" across; they sit on a long floral tube with the ovary at its base (attached to a leaf axil). The flowers have four yellow, heart-shaped petals, eight yellow stamens, and a cross-shaped stigma on a modest style; at the base of the flower are four sepals 3/8-3/4" long, reflected back along the floral tube as the flower develops with pairs of sepals attached along one edge until the flower opens. The sepals are often red-striped or red-tinged. Note the long, hairy floral tube below the sepals, the reddish stem, and glandular hairs with red bases on the ovary. The seed capsule develops from the ovary; it is up to 1.75" long, cylindrical but tapering slightly towards the tip, with four tiny lobes at the tip. — — — There are (at least) three evening primroses in Jackson Park — common evening primrose (Oenothera biennis), northern evening primrose (Oenothera parviflora), and hairy evening primrose (Oenothera villosa). They are easily confused with each other. (Judging by the conflicts between the accounts on Illinois Wildflowers [https://www.illinoiswildflowers.info/], Minnesota Wildflowers [https://www.minnesotawildflowers.info/], and Missouri Plants [http://www.missouriplants.com/], even the experts are confused.) All three species have yellow flowers with an X-shaped stigma, reflected sepals, petals with a small notch in their apex, stems that may be red or green, and lance-shaped leaves with short, widely-spaced teeth. Northern evening primrose is distinguished by (1) the presence of a knob or ridge at the tip of each sepal which the other two species lack (2) a seed capsule with four robust, flaring lobes with rounded tips at the apex of the seed capsule, and (3) a stigma that is very robust and almost fills the opening of the flower. Northern evening primrose has green to yellowish-green sepals. Hairy evening primrose (often but not always) has (1) striped or reddish-tinged sepals (check the buds where this character is often easier to see), (2) flowers that turn orange as they begin to senesce (although the reliability of this character is uncertain), and (3) always has glandular hairs covering the ovary that have "pimple-like" bright-red bases; these red pimples may also occur on the stems. All three species are somewhat similar to prairie sundrop (O. pilosella); the latter has very hairy stems and leaves, and translucent lines that radiate from the base of the petals in the flowers.](https://jacksonparkwildflowers.info/wp-content/uploads/2025/03/Oenothera_villosa-1t-300x300.jpg)
