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Large-flowered trillium's single flower sits on a 1-3" long flower stalk; the flower is 3-4" across and leans to one side (but doesn't droop). The flower consists of three green, lance-shaped sepals with rounded bases, three white (pinkish in older flowers) petals with wavy edges, six stamens with golden yellow anthers, a central white ovary, and three styles fused basally but with separated, dull-yellow stigmas. The fruit is a six-angled, globose capsule, about 1/2" long, that is initially green but darkens as it matures; it splits open to release the seeds. The seeds carry an attached oily, fleshy food reward (an elaiosome) and are dispersed by ants which inadvertently deposit the seeds in their nests to take advantage of the reward. Large-flowered trillium prefers shady environments like woodlands and is slow to mature — it can take several years for a sprout to produce flowers.

Trillium

Trillium grandiflorum

large-flowered trillium [Blooms: Apr-May]

The apical flowers of prairie trillium have three green sepals that are reflexed so they point down, three maroon petals, six maroon stamens with dark purple anthers incurved toward the center of the flower, and three purple styles with divergent stigmas hidden from view by the stamens. The fruit is a green, six-angled, ovoid berry 1" long; the seeds have an oily food appendage (an elaiosome) that encourages ants to carry them away. Despite the name, prairie trillium does not occur on prairies; it is a shade-loving, woodland plant like the other local trillium, large-flowered trillium (Trillium grandiflorum). Prairie trillium is superficially similar to toadshade (Trillium sessile), but in toadshade the sepals are spread (perpendicular to the stalk), not recurved (parallel to the stalk, pointed down). In addition, the leaves of toadshade are sessile (no petiole) and rarely have the troutlily-like mottling on the leaves that is seen on prairie trillium. The stamens of toadshade are yellow, not maroon. Toadshade does occur in Illinois, but it is less common, and I have yet to see it in Jackson Park.

Trillium

Trillium recurvatum

prairie trillium [Blooms: Apr-May]

The stalk of narrow-leaved cattails terminates in two spikes of unisexual flowers separated by a distinct gap (at least 1/2", usually several inches). The upper (staminate) spike is narrowly cylindrical, up to 8" long and 1/2" across, light yellow or light brown, and densely covered with staminate flowers (each bearing a single quartet of pollen grains) and numerous hairs. The lower (pistillate) spike is also cylindrical, up to 12" long and 3/4" across, brown or reddish brown in color, and densely covered with pistillate flowers and numerous hairs. The fertile pistillate flowers consist of a 1 mm long bractlet and a 1 mm long stipe that includes the ovary and a single, slender style; both are surrounded by several hairs. (Flora of North America (http://dev.floranorthamerica.org/Typha_angustifolia) and the Jepson Herbarium (https://ucjeps.berkeley.edu/eflora/eflora_display.php?tid=47466) have nice illustrations.) The flowers are wind pollinated. Some pistillate flowers are infertile and lack ovaries. Fertilized flowers produce a naked seed attached by a stalk to a tuft of white fibers; the seeds are dispersed by the wind. Narrow-leaved cattail can grow on wet soil or in water up to a foot deep.

Typha

Typha angustifolia

narrow-leaved cattail [Blooms: Jun-Jul]

In common cattail, staminate (male) flowers are located in a spike on the top of the cattail less than a half inch above the spike of female flowers (typically the spikes abut). The upper (staminate) spike is narrowly cylindrical, up to 12" long and 3/4" across, yellow or light brown, and densely covered with staminate flowers (each bearing a single quartet of pollen grains) and numerous hairs. The lower (pistillate) spike is also cylindrical, up to 18" long and 1.5" across, greenish or blackish brown in color, and densely covered with pistillate flowers and numerous hairs. The staminate spike withers soon after the pollen is released; the pistillate spike persists through the autumn. The fertile pistillate flowers consist of a 4 mm long stipe that includes the ovary and a single, slender 2.5 mm long style with a flattened stigma; both are surrounded by several hairs. (The Jepson Herbarium (https://ucjeps.berkeley.edu/eflora/eflora_display.php?tid=47466) has a nice illustration.) The flowers are wind pollinated. Some pistillate flowers are infertile and lack ovaries. Fertilized flowers produce a naked seed attached by a stalk to a tuft of white fibers; the seeds are dispersed by the wind. Common cattail can grow on wet soil or in water up to 1.5 feet deep. It often reproduces vegetatively from horizontal rhizomes buried in the soil/mud. The narrow-leaved cattail (Typha angustifolia) has narrower leaves (1/2"), smaller flower spikes (both in length and diameter) and the male flower spike is located a few inches above the female flower spike.

Typha

Typha latifolia

common cattail [Blooms: Jun-Aug]

Stinging nettle plants are dioecious (unisexual) with male (staminate) and female (pistillate) flowers on separate plants. Flowers occur in knobby clusters (panicles) that arise from the leaf axils. Neither pistillate nor staminate flowers have any petals. Staminate flowers have four light green, identical sepals cupping four short stamens. Pistillate flowers also have four sepals but two are narrowly lance-shaped, about a millimeter long, and free, alternating with two sepals that are 1.4-1.8 mm long and cupped around the ovary; styles are absent but the single stigma is long and feathery. The fruit is a flattened, oval to egg-shaped seed, less than 1/16" long.

Urtica

Urtica dioica

stinging nettle [Blooms: Jul-Aug]

Each side stem of yellow bellwort produces a single, elongate bell-shaped flower at its tip. The flowers droop on their stem-like pedicels (about 1 " long) such that they open downwards and are often partially hidden by the leaves. Each flower is 1-2" long and consists of six yellow tepals twisted along their length (often with greenish bases), six white stamens with elongated yellow anthers, and a pistil with a three-lobed stigma. The fruit is a three-section capsule that looks (vaguely) like three fat triangles with oval bases, stuck base to base to base. Seeds are freed when the capsule splits. Seed dispersal is added by the attraction of ants to a fleshy, nutrient-rich appendage (an elaiosome) on the seed; the ants carry the seed back to their nest where they eat the elaiosome and abandon the seed.

Uvularia

Uvularia grandiflora

yellow bellwort [Blooms: Apr-May]

The apex of the common mullein plant is a club-shaped flower spike 6-24" long, covered with scattered light-yellow flowers each 3/4" across. Individual flowers have five hairy, green sepals; five pale yellow petals (the lower two slightly larger than the upper three); five stamens with reddish-orange anthers; and a single green style with a heart- or disc-shaped stigma. The upper three stamens are shorter than the other two stamens and covered with white or yellow hairs; the lower two stamens are longer and largely hairless. The fruit is an oval, two-celled seed capsule, brown when mature, each cell filled with numerous (25-150) tiny orange to brown seeds. Common mullein is immediately recognizable and impossible to confuse with any other local plant.

Verbascum

Verbascum thapsus

common mullein [Blooms: Jun-Aug]

The top of the wingstem plant is a dome (a panicle) of coarse-looking flowerheads. Each flowerhead is 1-2" across, with 2-10 yellow, oblong, drooping petals, 1/2-1" long that are derived from the sterile ray florets. Some 40-60+ yellowish-green, tubular disk florets project in all directions in a hemisphere of widely-spaced, robust, bud-like blunt cylinders, the tips of the disk florets. When receptive to pollen, the disk florets bear a protruding style with a bifurcated, curling pair of stigmas (likened by one source to a "pince-nez" — a pair of eyeglasses on a handle); the five stamens and their flattened, black anthers are appressed to the body of the style. The fruit is a teardrop-shaped seed with a pair of thin wings and two short, robust, spiky awns. Immature seeds are green; the seeds (and seedheads) turn brown when mature.

Verbesina

Verbesina alternifolia

wingstem [Blooms: Jul-Oct]

Wayfaring tree flowers occur in flat-topped cymes about 4" across; they are cream-colored with five petals, five stamens with white filaments and tan anthers, and a green ovary with a short (1 mm) style with a large, lobed stigma. The fruit is an oval, flattened drupe, initially red, turning black with maturity, and containing a single seed. They are not toxic but can induce vomiting it eaten when unripe. Wayfaring tree is said to be so named because it grows close to paths (and presumably other natural edges).

Viburnum

Viburnum lantana

wayfaring tree [Blooms: Apr-May]

Nannyberry flowers occur in 2-5" wide, dome-shaped panicles at the tips of young shoots. Individual flowers are 1/4" across, consisting of a light green calyx with five triangular teeth; a creamy white corolla with five oval lobes; five stamens with white filaments and yellow anthers that protrude well past the edges of the corolla lobes; and a light green or yellow pistil with a short style. The fruit is a 6-10 mm long drupe, initially green but turning dark blue-violet when ripe, containing a single seed. Nannyberry is similar to blackhaw (Viburnum prunifolium, also present in Jackson Park) — both have white flowers and blue-black fruits that are drupes — but nannyberry leaves have a long, narrow tip on their leaves in contrast to the rather blunt tips on blackhaw leaves.

Viburnum

Viburnum lentago

nannyberry [Blooms: Apr-May]

In the spring, American cranberry shrubs produce flattened panicles of flowers 2-5" across that bear two types of flowers — around the edges are five-petaled, white flowers about 1" across that are sterile (lacking both pistils and stamens), while in the center of the cluster are five-petaled, cream-colored flowers 1/4" across that are fertile (with both a pistil and five long stamens). By late summer, the flowers are replaced by bright red "cranberries" (actually drupes with a single seed) 1/4-1/2" across that prove very popular with birds (which distribute the seeds) and our local coyotes (whose scat is full of the seeds). After removing the seed, the tart flesh (high in vitamin C) can be eaten raw or cooked. In the fall, the leaves also turn red. Viburnum opulus var. americanum is said to be distinct from Viburnum trilobum (aka, highbush cranberry) by Illinois Wildflowers (www.illinoiswildflowers.info) and Minnesota Wildflowers (www.minnesotawildflowers.info), but plants.usda.gov insists that the latter is a synonym of the former. Worldfloraonline.org only adds to the confusion by insisting that the two names are BOTH junior synonyms of Viburnum opulus. ITIS.gov says that V. opulus is the correct name for the European species (now introduced to the U.S.) and Viburnum opulus var. americanum is the correct name for the American species. The easiest way to distinguish the two forms (if you care to) is to carefully inspect the nectary glands (extrafloral nectaries) on the petioles. In V. opulus americanum, the extrafloral nectaries are oval, with a concave top and a distinct rim; in Viburnum trilobum, the extrafloral nectaries are round in plan view with a convex top and no rim. Although rare in most of Illinois, guelder-rose (V. opulus americanum) is quite common around the lagoons of Jackson Park.

Viburnum

Viburnum opulus

American cranberry [Blooms: Apr-May]

Cymes of flowers arise from the leaf axils of blackhaw; the cymes are heavily branched but are sessile on the leaf axil. Individual flowers are 1/4" across with a 2 mm long calyx tube with five apical lobes that are shorter than the petals; a tubular corolla with five white, rounded lobes (the "petals"); five stamens with white filaments and yellow anthers extending outside the corolla lobes; and a small (less than 1 mm long), cream-colored pistil. The fruit is a fleshy drupe (like a plum or cherry) about 8 mm long containing a single brown nutlet that is flat on one side; the fruit becomes blue-black as it matures. The fruits are edible and popular with birds. Blackhaw is similar to nannyberry (Viburnum lentago, also found in Jackson Park) — both have white flowers and blue-black fruits that are drupes — but nannyberry leaves have a long, narrow tip on their leaves in contrast to the rather blunt tip and reddish petioles on blackhaw leaves. The stamens in blackhaw flowers are slightly shorter than those in nannyberry flowers but the difference is subtle; leaf form and petiole color are better differentiators.

Viburnum

Viburnum prunifolium

blackhaw [Blooms: May-June]

Many of the upper shoots of downy arrowwood end in 1.5-3" wide, dome-shaped panicles of flowers with slightly fuzzy, light green branches. Individual flowers are 1/4" across with a short, light green calyx with five short, broad teeth; a white tubular corolla with five rounded, spreading lobes; five stamens with white filaments and tan anthers that extend outside the corolla; and a single white style. The flowers have an unpleasant odor described by various sources as "rotting flesh." The fruit is a small (1/4-1/2") drupe (a stone fruit like cherries or a plum) 8 mm long and dark blue-violet to black at maturity. Downy arrowwood can be distinguished from other Viburnum spp. by its (1) its small size (less than six feet tall), (2) the relatively few teeth along the leaf margins (5-10 per side), (3) the short (about 1/4") leaf petioles, (4) the small, narrow stipules at the petiole bases, and (5) its flowers' unpleasant odor. The common name derives from the fact that the straight stems of the various arrowwoods was preferred by indigenous Americans for arrow shafts.

Viburnum

Viburnum rafinesquianum

downy arrowwood [Blooms: Aug–Sep]

Riverbank grape's flowers develop in 5" long, drooping, cylindrical or pyramidal panicles opposite the leaves of new branches (but skipping every third leaf). The flowers are greenish-yellow and are best described as skeletal; they may be either unisexual or (rarely) perfect (both male and female reproductive organs functional). The male and female unisexual flowers usually both occur on the same plant, either in separate or mixed clusters; both male and female flowers are 1/8" across or less, with a minute calyx and five green to yellowish petals that quickly fall off. Male flowers have five very long, pale, erect stamens around a button-like center; female flowers have an ovary with a short style and five short, sterile stamens, often contorted. The fruit are .... grapes — a round berry, 1/4-1/2" across containing 1-4 seeds, initially green but turning blue-black as it matures, often with a white bloom on the skin. You know, grapes. The fruit of riverside grape is often sour until after a frost, then becomes more "sweet/tart" to the tongue. Riverside grape is subject to parasitism by the grape filbert gall midge (Ampelomyia vitiscoryloides), an insect whose larvae produce large, very hairy, conical galls at leaf nodes. This gall midge is restricted to the eastern half of North America; it was first described by Alpheus Spring Packard, Jr. in 1869.

Vitis

Vitis riparia

riverbank grape [Blooms: [May-Jun]

Common cocklebur produces a spike-like raceme of compound flowerheads, shorter than the leaf petioles, that arises from the axils of each upper leaf and from the tip of the central stem. Each raceme produces exclusively male flowerheads in its upper parts and exclusively female flowerheads in the lower regions. The male flowerheads are about 1/4-3/4" across, elevated on a short pedicel on top of 1-3 series of white floral bracts. Each of the approximately 20 tiny, brownish-green staminate florets (nominally disk florets) consist of a tubular 1-2 mm long corolla; a minute, non-functional ovary; and five stamens with fused filaments (like a tree trunk) but with their white anthers free, positioned in a ring like a little palm tree or, possibly, a hand. The staminate florets quickly shed their pollen and drop off. The female flowerheads (also nominally disk florets) are 3/4-1.5" long and two-thirds as wide, stalkless, each (usually) containing two female florets nearly enclosed by floral bracts covered with curly white hairs that have a bur-like appearance from the dense, hooked prickles covering their surface. At the tip of the flowerhead are two spines, longer and much stouter than the prickles, each with a hole from which protrudes the divided style of a female floret. There is no corolla in the female florets. The florets are wind pollinated, and their styles quickly wither away after fertilization. Each female floret produces a single, oblong, dark gray to black seed 3/8-1/2" long, tapering to a point at each end and covered by dark membranes, all buried in the bur-like bract. The female flowerhead stiffens through the autumn, turns brown, and overwinters. The seeds are highly poisonous, containing toxic diterpene glycosides. One of the contained seeds germinates the following year; the other delays germination for two years. Common cocklebur prefers wet, sandy soils.

Xanthium

Xanthium strumarium

common cocklebur [Blooms: Sep-?]

The tips of the upper stems of heartleaf alexanders produce 2-3" wide compound umbels of 7-15 umbellets, each with 10-20 tiny yellow flowers. The central flower of each umbellet is sessile; the rest have short pedicels (a diagnostic character of the genus). Each 1/8" wide flower consists of five yellow petals that remain curled, five stamens with yellow anthers projecting through the spaces between the petals, and two styles. The fruits are a ribbed capsule with a flat top from which the remnants of the styles project. Heartleaf alexanders is restricted to northeastern Illinois, where it is uncommon; it is usually found in drier habitats than its much more common relative, golden alexanders (Zizia aurea).

Zizia

Zizia aptera

heartleaf alexanders [Blooms: Apr/May–?]

The tips of the upper stems of golden alexanders produce flat to slightly rounded compound umbels 2-3" across with about a dozen umbellets of tiny, golden-yellow flowers. Each umbellet has about 20 flowers; the central flower is sessile (or nearly sessile) while all the other blooms in the umbellet are on short stalks (pedicels), a character of the genus. Each 1/8" wide flower consists of an insignificant calyx, five yellow petals that are persistently incurled, five stamens with yellow anthers projecting through the spaces between the petals, and two styles. The fruits are a ribbed (not winged) 2 mm long capsule with a flat top from which the remnants of the two styles project. Golden alexanders prefers full to partial sun and is tolerant of a range of habitat quality. The yellow-flowered variety of meadow parsnip (Thaspium trifoliatum var. aureum) is very similar to golden alexanders but has basal leaves that are simple rather than trifoliate compound, but I have not seen meadow parsnip in Jackson Park. John Hilty (www.illinoiswildflowers.info) warns that "Golden Alexanders should not be confused with Pastinaca sativa (Wild Parsnip), which is a weedy Eurasian biennial. The latter is taller, blooms later, and has more leaflets in each compound leaf." Wild parsnip DOES occur in Jackson Park, but it is relatively rare, twice the size of golden alexanders, has robust, strongly ribbed stems, and a large, flattened, winged fruit. Heartleaf alexanders (Zizia aptera) is also very similar, but only the uppermost leaves are compound; all the lower and middle leaves are simple and heart-shaped.

Zizia

Zizia aurea

golden alexanders [Blooms: Apr/May–?]