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Purple meadow-rue is a native perennial that is listed as "vulnerable" in Illinois. The species is found in most of the lower 48 states in the U.S. but is notably absent from states on the Atlantic and Pacific coasts (however, it is present along the Gulf of Mexico). Purple meadow-rue is dioecious — some plants produce only staminate (male) flowers, each with around a dozen stamens; other plants bear only pistillate (female) flowers, each with about ten pistils. The male flowers are the showier of the two. Petals are absent (the plant is wind-pollinated). Both the narrow, whitish-grey, thread-like pistils and the yellow-green stamens (with pale yellow anthers) droop from 4-5 light green sepals (that may drop off). Fertilized female flowers develop a single, spindle-shaped fruit that is 1/4" long, elongated, sharply pointed with a remnant style, 4-6 raised ribs, and slightly concave along one side. The fruit is initially green, turning brown with maturity. Early meadow-rue (Thalictrum dioicum) is similar to purple meadow-rue but has light purplish-green stems and leaves with 3-9 (typically five) rounded lobes and long petioles on the upper leaves.

Thalictrum

Thalictrum dasycarpum

purple meadow-rue [Blooms: May-Jun]

Early meadow-rue's main stem terminates in a panicle up to a foot long and wide; each branch in the panicle has a corymb of about five flowers at its tip. Early meadow-rue is dioecious — some plants produce only staminate (male) flowers, each with 10+ stamens; other plants bear only pistillate (female) flowers, each with up to 15 pistils. Both male and female flowers droop from their stems, but the male flowers have a longer droop than do the female flowers. Both male and female flowers are about 1/2" long and 1/4" across with 4-5 pale green sepals with white margins (which often fall off). Neither male nor female flowers have petals; pollination is via the wind. Fertilized female flowers produce ellipsoid, dry seeds that are pointed at both ends; the surface bears strong ribbing. Early meadow-rue prefers shady woodlands. It has more lobes on its leaves (usually 3-5), and the lobes are more rounded than purple meadow-rue (Thalictrum dasycarpum), the other Thalictrum species in Jackson Park, does (2-3 pointed lobes).

Thalictrum

Thalictrum dioicum

early meadow-rue [Blooms: Apr-May]

Erect hedge-parsley's main stem and side stems give rise to umbels 1.5-2" across of 10-20 umbellets of flowers. Each flower is about 1/8" across consisting of five white petals (often markedly unequal in size), and a creamy white center surrounded by five pink to white stamens. The fruit is an 1/8" long oval covered in hooked hairs to aid in bur-like dispersal; it contains two seeds. (The fruit of common hedge-parsley (Torilis arvensis) has straight or only slightly curved hairs, not hooked, but the hairs are barbed.)

Torilis

Torilis japonica

erect hedge-parsley [Blooms: Jun-Jul]

Fuzzy flower stalks about 1" long arise from the leaf axils of low hop clover. The flowerheads are roughly spherical, lemon yellow, and about 1/2" across comprised of 20-40 pea-like florets with five petals; each floret is about 4 mm long. The upper petal (the standard) on each floret is conspicuously grooved and forms a hood over the two petals forming the keel; two lateral petals flank the keel. Fertilized florets form tiny seedpods that are shorter than the keel. The flower color and foliage of low hop clover are very similar to black medic (Medicago lupulina), but the standard on black medic is not grooved and sits perpendicular to the keel.

Trifolium

Trifolium campestre

low hop clover [Blooms: May-Aug]

Alsike clover is an exotic perennial native to Europe. Despite the scientific name ("hybridus"), it is not a hybrid. (Linnaeus thought it was a hybrid of red and white clover. It isn't.) The plant is 6-24" tall, with occasional branches along the stems. One- to three-inch-long flower stalks arising from the leaf axils terminate in a flowerhead 1/2-3/4" across; florets tend to be white on the top of the flowerhead and reddish-purple to pink on the bottom, but may all be reddish. Individual florets are 1/4" long with five petals in a pea-like bloom. They have a sharply pointed standard (the major vertical petal, perpendicular to the keel), two petals that make up the keel, and two lateral petals that flank the keel. Ten stamens and a single pistil are hidden inside the keel. Unlike red clover (T. pratense) and white clover (T. repens), there are no pale chevron-shaped markings on the leaflets.

Trifolium

Trifolium hybridum

alsike clover [Blooms: May-Jul]

The upper stems of red clover end in 3/4-1" wide pinkish-purple (not actually red) flowerheads, usually with 1-3 leaflets or a compound leaf and some pointed bracts immediately below the bloom. The florets are tubular modification of a typical pea-like flower with five petals; the upper petal (the standard) is slightly longer than the other four petals and has a slightly pointed tip. Red clover is easily identified by the pinkish-purple flowerheads, the leaflets inserted just below the flowerheads, and the white chevrons on the leaflets.

Trifolium

Trifolium pratense

red clover [Blooms: Apr-Jun]

White clover's hairless, leafless flower stalks are up to 3" long. They emerge directly from the horizontal stems and grow perpendicular to the ground, producing a single globular 3/4" wide flowerhead at their tips. The flowerheads are all white, consisting of 20-50 pea-like florets. The florets are tubular, consisting of a green calyx with five long, narrow teeth, and five white or pinkish-white petals (a standard, two petals making the keel, and a pair of lateral petals flanking the keel). White clover is similar to alsike clover (T. hybridum), but that species usually has some pink in the flowerheads and never has white chevrons on the leaflets.

Trifolium

Trifolium repens

white clover [Blooms: May-Jun]

Clusters of lowbush blueberry's flowers develop on the previous year's twigs. The flowers are about 1/4" long, urn- or bell-shaped, nodding, white, and waxy. The flowers contain a short, green calyx with five teeth; a short, tubular corolla that is pinkish white or white with five triangular, recurved lobes around its margin; ten stamens with reddish brown anthers recessed behind the margin of the corolla and clustered around the style; and a single style whose stigma extends right to the margin of the corolla. Fertilized flowers are replaced by approximately 8 mm diameter berries, initially green but turning dark blue with a whitish bloom as they mature. (I.e., they look — and taste — like blueberries.)

Vaccinium

Vaccinium angustifolium

lowbush blueberry [Blooms: May]

The apex of the common mullein plant is a club-shaped flower spike 6-24" long, covered with scattered light-yellow flowers each 3/4" across. Individual flowers have five hairy, green sepals; five pale yellow petals (the lower two slightly larger than the upper three); five stamens with reddish-orange anthers; and a single green style with a heart- or disc-shaped stigma. The upper three stamens are shorter than the other two stamens and covered with white or yellow hairs; the lower two stamens are longer and largely hairless. The fruit is an oval, two-celled seed capsule, brown when mature, each cell filled with numerous (25-150) tiny orange to brown seeds. Common mullein is immediately recognizable and impossible to confuse with any other local plant.

Verbascum

Verbascum thapsus

common mullein [Blooms: Jun-Aug]

The tip of a smooth ironweed central stem bears a cluster (a corymb) up to 4" in diameter of magenta composite flowerheads that is quite densely packed in a flat or (sometimes) domed inflorescence. The branches of the inflorescence (but not the stem) may be lightly fuzzy with small hairs. Each flowerhead is 3/4" across and contains 10-30 magenta disk florets, each with five lobes and a very prominent, divided style with curled tips (the stigmas) and five stamens appressed to the base of the style; ray florets are completely absent. The bracts that cover the underside of the flowerheads are green or purplish brown, flattened, appressed, and rounded at their tips, imbricated like fish scales with white, cobwebby hairs around their edges. The fruit is a naked seed with an attached tuft of white hairs (actually, scales). To sort out the Jackson Park ironweeds: (1) if both leaves and stem are densely covered with fine white hairs, the leaves are about 3-4 times longer than wide, and each flowerhead has more than 30 disk florets, it's Missouri ironweed (V. missurica). (2) If both leaves and stems are hairless, there are less than 30 disk florets, and the leaves are narrow (typically a half inch or less; about 6-10+ times longer than wide), it's smooth ironweed (V. fasiculata). (3) If the stems and leaves have only short hairs, there are less than 30 disk florets, and the leaves are about 2" wide and 4-6 times longer than wide, it's tall ironweed (V. gigantea).

Vernonia

Vernonia fasciculata

smooth ironweed [Blooms: Jun-Sep]

Tall ironweed's inflorescence is a flat-topped panicle arising from the tip of the central stem; it is 6-16" across, open and spreading. There are 10-30 disk florets in a tall ironweed flowerhead. The base of the flowerhead is covered by several series of dark colored, broadly oval appressed bracts (phyllaries), often with fine hairs around their edges. The disk florets have tubular, trumpet-like corollas with five recurved lobes; ray florets are absent. Tall ironweed has a much more "open" composite flowerhead than the other ironweeds in Jackson Park, with the trumpet bases of the disk florets easily visible. The fruits are naked, gray or brown, bullet-shaped seeds with a tuft of both short and long tawny or pale purple hairs. To sort out the Jackson Park ironweeds: (1) if both leaves and stem are densely covered with fine white hairs, the leaves are about 3-4 times longer than wide, and each flowerhead has more than 30 disk florets, it's Missouri ironweed (V. missurica). (2) If both leaves and stems are hairless, there are less than 30 disk florets, and the leaves are narrow (typically a half inch or less; about 6-10+ times longer than wide), it's smooth ironweed (V. fasiculata). (3) If the stems and leaves have only short hairs, there are less than 30 disk florets, and the leaves are about 2" wide and 4-6 times longer than wide, it's tall ironweed (V. gigantea).

Vernonia

Vernonia gigantea

tall ironweed [Blooms: Jul-?]

At the tip of a Missouri ironweed stem, a corymb of composite flowerheads (the inflorescence) is produced. The stems of the inflorescence are also covered in fine white hairs and usually reddish brown in color. Individual flowerheads are magenta and rayless; ray florets are completely absent. Flowerheads contain 32-60 disk florets, each 1/2-3/4" across, which have rounded or broadly angled bracts (phyllaries) with sharp tips, dull green or reddish brown in color, that are layered like fish scales around the base. To sort out the Jackson Park ironweeds: (1) if both leaves and stem are densely covered with fine white hairs, the leaves are about 3-4 times longer than wide, and each flowerhead has more than 30 disk florets, it's Missouri ironweed (V. missurica). (2) If both leaves and stems are hairless, there are less than 30 disk florets, and the leaves are narrow (typically a half inch or less; about 6-10+ times longer than wide), it's smooth ironweed (V. fasiculata). (3) If the stems and leaves have only short hairs, there are less than 30 disk florets, and the leaves are about 2" wide and 4-6 times longer than wide, it's tall ironweed (V. gigantea).

Vernonia

Vernonia missurica

Missouri ironweed [Blooms: Jul-Oct]

Wild pansy's flower stalks are long, hairless, and bend like a shepherd's crook just below the base of the flower so the flower axis is approximately horizontal. Individual flowers have five 7-15 mm long, lance-shaped sepals that are longer than the petals and are sharply pointed at their tips; a 4-12 mm long corolla with five white or pale cream-colored petals, a yellow throat with a few dark purple lines, the lateral petals bearded on their upper surface, and a 1-1.5 mm long spur; two stamens not visible externally; and a single style enlarged into a hollow, globose stigma. The fruit is a spherical or ellipsoidal three-valved capsule, 5-10 mm long containing 6-75, 1.5 mm long, brown seeds.

Viola

Viola arvensis

wild pansy [Blooms: Jun-?]

Yellow violet is a native perennial, the only yellow violet species in Jackson Park (indeed, in all of Illinois). One or more stems arise from an axil of a stem leaf; they are up to 6" tall with a single 3/4" long flower at their tips. Each flower consists of five light green sepals, five yellow petals, five stamens, and a single style; the stamens and style cannot be seen from the exterior. On the lower petal, the flowers have purple lines (nectar guides) and a spur reduced to a small pouch. The two lower, lateral petals have small tufts of hair that obscure the throat of the flower and hide the stamens and style; the two upper, lateral petals are beardless and have only faint lines (nectar guides), if any. The fruit of this species is a bullet-shaped (hairless in the Jackson Park forms) capsule nestled in the calyx with the remnant of the style at its tip, initially green but turning brown as it matures; it ultimately splits along three suture lines to fling the seeds away from the plant.

Viola

Viola pubescens

yellow violet [Blooms: Apr-May]

Riverbank grape's flowers develop in 5" long, drooping, cylindrical or pyramidal panicles opposite the leaves of new branches (but skipping every third leaf). The flowers are greenish-yellow and are best described as skeletal; they may be either unisexual or (rarely) perfect (both male and female reproductive organs functional). The male and female unisexual flowers usually both occur on the same plant, either in separate or mixed clusters; both male and female flowers are 1/8" across or less, with a minute calyx and five green to yellowish petals that quickly fall off. Male flowers have five very long, pale, erect stamens around a button-like center; female flowers have an ovary with a short style and five short, sterile stamens, often contorted. The fruit are .... grapes — a round berry, 1/4-1/2" across containing 1-4 seeds, initially green but turning blue-black as it matures, often with a white bloom on the skin. You know, grapes. The fruit of riverside grape is often sour until after a frost, then becomes more "sweet/tart" to the tongue. Riverside grape is subject to parasitism by the grape filbert gall midge (Ampelomyia vitiscoryloides), an insect whose larvae produce large, very hairy, conical galls at leaf nodes. This gall midge is restricted to the eastern half of North America; it was first described by Alpheus Spring Packard, Jr. in 1869.

Vitis

Vitis riparia

riverbank grape [Blooms: [May-Jun]

American wisteria's flowers occur in showy terminal racemes. The flowers have 1/2" long pedicels that are densely covered with stiff hairs and some club-shaped, stalked glands. Individual flowers consist of a 1/4-3/8"white to pale green, hairy calyx and 15-20 mm long bluish-purple, pea-like, hairless flowers. The expanded portion of the banner (or standard) is bent backwards so it is perpendicular to the axis of the rest of the flower; the two lateral petals cover the keel like a hood. There are 10 stamens with tiny anthers, an ovary encircled by a nectar gland, and a single style hidden within the keel; nine of the 10 stamens are fused to each other, the tenth is free. The fruits are 2.75-4" long pea-like, hairless green pods containing 4-8 kidney-shaped or oval seeds, each about 8 mm wide; the pods are brown and the seeds are black when mature. The seeds contain an alpha-amino acid called canavanine which is structurally similar to L-arginine and the pods contain a saponin called wisterin; neither is a good thing to ingest. I've only seen American wisteria growing on the fence west of Japanese Garden. Periodically, the Park District cuts it down, I suspect because they fear for the fence's structural integrity (it's a heavy plant), but I wish a better solution could be found given American wisteria's rarity.

Wisteria

Wisteria frutescens

American wisteria [Blooms: May-Oct]

The tips of the upper stems of heartleaf alexanders produce 2-3" wide compound umbels of 7-15 umbellets, each with 10-20 tiny yellow flowers. The central flower of each umbellet is sessile; the rest have short pedicels (a diagnostic character of the genus). Each 1/8" wide flower consists of five yellow petals that remain curled, five stamens with yellow anthers projecting through the spaces between the petals, and two styles. The fruits are a ribbed capsule with a flat top from which the remnants of the styles project. Heartleaf alexanders is restricted to northeastern Illinois, where it is uncommon; it is usually found in drier habitats than its much more common relative, golden alexanders (Zizia aurea).

Zizia

Zizia aptera

heartleaf alexanders [Blooms: Apr/May–?]

The tips of the upper stems of golden alexanders produce flat to slightly rounded compound umbels 2-3" across with about a dozen umbellets of tiny, golden-yellow flowers. Each umbellet has about 20 flowers; the central flower is sessile (or nearly sessile) while all the other blooms in the umbellet are on short stalks (pedicels), a character of the genus. Each 1/8" wide flower consists of an insignificant calyx, five yellow petals that are persistently incurled, five stamens with yellow anthers projecting through the spaces between the petals, and two styles. The fruits are a ribbed (not winged) 2 mm long capsule with a flat top from which the remnants of the two styles project. Golden alexanders prefers full to partial sun and is tolerant of a range of habitat quality. The yellow-flowered variety of meadow parsnip (Thaspium trifoliatum var. aureum) is very similar to golden alexanders but has basal leaves that are simple rather than trifoliate compound, but I have not seen meadow parsnip in Jackson Park. John Hilty (www.illinoiswildflowers.info) warns that "Golden Alexanders should not be confused with Pastinaca sativa (Wild Parsnip), which is a weedy Eurasian biennial. The latter is taller, blooms later, and has more leaflets in each compound leaf." Wild parsnip DOES occur in Jackson Park, but it is relatively rare, twice the size of golden alexanders, has robust, strongly ribbed stems, and a large, flattened, winged fruit. Heartleaf alexanders (Zizia aptera) is also very similar, but only the uppermost leaves are compound; all the lower and middle leaves are simple and heart-shaped.

Zizia

Zizia aurea

golden alexanders [Blooms: Apr/May–?]