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At the tip of the culm, dark green bulrush produces an inflorescence comprised of one to three (rarely four or five) compound umbels of spikelets; beneath the inflorescence are 3+ leafy bracts (similar to the leaves) that are up to 1' long and 1/2" wide. At least one of the bracts is always longer than the inflorescence. The umbels generate 1/4-4" long, straight branches in all directions that terminate in umbelletes of 5-20 spikelets that rest on slender, leafy bractlets; the spikelets may be sessile or may rest on 1/2" long branchlets. The branches and branchlets are green, straight, and stiff. Spikelets are less than 1/4" long (3-6 mm), ovoid or egg-shaped, yellowish green initially but darkening to brown or black later; the spikelets are an assemblage of numerous spirally arranged, overlapping scales and their florets. The florets each have a single subtending scale, three stamens, and an ovary with a tripartite style; they are wind pollinated. Bulblets may form on one or more branches and sprout late in the season. The umbels may be visually dense and complex and take some study to sort out.

Scirpus

Scirpus atrovirens

dark green bulrush [Blooms: May-Jul]

Early figwort (aka, lanceleaf figwort) is difficult to distinguish from late- (or Maryland-) figwort (S. marilandica) except by blooming date (before July = early, after July = late, July = could be either) and details of the flowers (especially the sterile stamen). The inflorescence (a panicle) is constructed from short, spreading, sparse branches covered in glandular hairs, bearing a 3/8" long, shiny tubular flower whose outside surfaces are reddish-purple to reddish-green and inner surfaces are reddish brown (upper and lateral lobes) or greenish yellow (lower lobe). Four yellow, fertile stamens and a single blunt, slender style hug the floor of the flower, the style protruding further than the fertile stamens. A single sterile stamen (a "staminode") is located along the upper interior of the flower. The sterile stamens are yellow or green with a broad tip in early figwort (S. lanceolata); they are reddish-brown to purple and narrow in late figwort (S. marilandica). The fruit is a teardrop-shaped capsule, dark brown when mature, that splits in two longitudinally to release the numerous tiny seeds.

Scrophularia

Scrophularia lanceolata

early figwort [Blooms: May-Jul]

Late figwort's inflorescence (a panicle) gives rise to short, spreading, sparse branches covered in glandular hairs, bearing a 3/8" long, shiny tubular flower whose outside surfaces are reddish-purple to reddish-green and inner surfaces are reddish brown (upper and lateral lobes) or greenish yellow (lower lobe). Four yellow, fertile stamens and a single blunt, slender style hug the floor of the flower, the style protruding further than the fertile stamens. A single sterile stamen (a "staminode") is located along the upper interior of the flower. The sterile stamens are yellow or green with a broad tip in early figwort (S. lanceolata); they are reddish-brown to purple and narrow in late figwort (S. marilandica). The fruit is a teardrop-shaped capsule, dark brown when mature, that splits in two longitudinally to release the numerous tiny seeds.

Scrophularia

Scrophularia marilandica

late figwort [Blooms: Jul-Aug]

Mad-dog skullcap flower racemes may arise at the tips of the stems or from the axils of leaves. Each raceme is up to 6" long, containing 6-7 pairs of flowers; each flower has a short, basal leafy bract. Flowers are up to 8 mm long with a short, slender pedicel, a light green tubular calyx with two shallow lobes (the upper lobe with a dish-like projection), and a pale blue, lavender, or white, densely hairy, tubular corolla 5-8 mm long with a short, hood-like, upper lip and a slightly flared, three-lobed lower lip. Hidden inside the upper lip of the corolla are four stamens and a single bifurcated style. After fertilization, the calyx swells into a green capsule with includes the dish-like projection mentioned above, giving the capsule a unique appearance; inside is a cluster of four nutlets. The mouth of the calyx is visible as a thin slit beneath the projection.

Scutellaria

Scutellaria lateriflora

mad-dog skullcap [Blooms: Jul-Aug]

Crown vetch's flower stalks are about 6" long with a 1" wide cluster (an umbel) of 10-25 pea-like flowers on short (1/4") pedicels, all attached to the tip of the flower stalk. Each flower is 1/2" long and 1/4" wide, with a short green calyx bearing five pointed lobes and a typical pea-like corolla, bicolored in pink and white. The five petals are modified as a vertical standard (often pinker than the other petals, which are white), two lateral wings, and a pair of abutting petals forming a medial keel perpendicular to the standard. Within the keel are 10 stamens (one free and nine fused to each other, all with small yellow anthers) and a slender style, curved upwards, with a tiny terminal stigma. The fruit is a 1/2-2" long, narrow seedpod (loment) with 1-7 slightly constricted segments and a long terminal "beak" (the remains of the pistil); the pods remain attached to the flower stalk, forming a cluster often likened to "fingers". The tight flower clusters with showy, bicolored flowers are characteristic.

Securigera

Securigera varia

crown vetch [Blooms: Jun-Sep]

Biting stonecrop (aka, mossy stonecrop) is an exotic succulent that is native to Greenland and western Europe but naturalized across most of the U.S. and the lower Canadian provinces. It is a mat-forming plant with creeping stems that roots at nodes; the stems produce short, erect vegetative and flowering stems. The leaves are alternate, sessile, and densely imbricated on the erect stems; they are green or yellowish-green, with old, white leaves persisting. The succulent-like leaves are 4-5 mm long, 2.5-3 mm across, and 1.5 mm thick, convex on both the upper and underside and blunt-tipped. The inflorescence arises from the tip of one of the erect stems; it is about 5/8" (1.5 cm) long, 1/4-1.25" (5-30 mm) across, with 1-3 branches, each with 1-4 sessile (or on 1 mm long pedicels) flowers with a basal bract. The sepals closely resemble the leaves. The flowers have five bright yellow petals, typically 10 stamens with yellow anthers, and 4-5 yellow pistils, each with a single style and a pinhead-shaped stigma. The fruit is a five-pointed yellowish star of united, inflated follicles filled with numerous seeds. Biting stonecrop is not a good competitor; it grows best in thin, dry soils like cliffs, shingle, and beaches. However, it is tough — it stores water in its leaves and reportedly can go six months without external water.

Sedum

Sedum acre

biting stonecrop [Blooms: Jun]

The tips of the upper stems of common groundsel produce dense clusters of flowerheads on short peduncles; inflorescences can also arise from leaf axils. Common groundsel flowerheads are cylindrical, 8 mm long and 4 mm across, with black-tipped, green bracts around the outside at both the base and the apex and 13-80 yellow disc florets peeking out the end; mature flowerheads are easily mistaken for flower buds. The disc florets have tubular yellow corollas, five stamens fused to the upper part of the corolla with yellow-brown anthers, and a bifurcated style with purplish stigmas. There are no ray florets. This is an easy plant to overlook. The flowerheads are twice as long as wide. The seedhead looks like a compact dandelion tuft with seeds on parachutes of light brown hairs.

Senecio

Senecio vulgaris

common groundsel [Blooms: May-Jul]

The five petals and five sepals of Maryland senna's 3/4" flowers are both yellow. The single pistil has many spreading white hairs and arches upward. The anthers of the 10 stamens are a striking dark brown (like partridge pea). The upper three stamens have short filaments and short anthers that produce only sterile pollen (as a pollinator reward). The middle four stamens have short filaments and long anthers while the lower three stamens have long filaments and long anthers; all seven of these stamens produce fertile pollen that is released by "buzz pollination." The flowers do not produce nectar. The fruit is a flattened seedpod 1/2" across and 3-4" long with 10-18 segments, each containing a single 1/4" flattened seed.

Senna

Senna marilandica

Maryland senna [Blooms: Jul]

Flower buds arise singly or in small groups on 1/2" long peduncles from the leaf axils. Because the flower peduncles are shorter (1/2") than the leaf petioles (1"), the flowers are often partly hidden among the leaves. Individual flowers are about 8 mm across. A flower contains five green, triangular sepals; five light yellow or light orange, rounded, floppy petals with pale lines that function as nectar guides and asymmetrical tips; and a central reproductive column (like mallows, to which prickly sida is related). That column consists of five styles fused along most of their lengths, each with a globose terminal stigma, and numerous stamens with yellow anthers. After fertilization, a circular seedpod forms from the five carpels; when mature, the seedpod turns brown and breaks into five segments, each containing a single reddish-brown, three-angled seed.

Sida

Sida spinosa

prickly sida [Blooms: Aug-Sep]

Evening campion is an exotic from Europe that grows to be 2-3' tall. It is nominally a night-blooming species, but I've observed it in bloom throughout the day in shady areas. Male and female plants are separate. The upper stems end in clusters of one or more 1-1.5" long flowers. The "bladder" (the calyx) is covered with a fuzz of fine hairs and has dark green or reddish-purple lines running from pole to pole (10 lines in male flowers, 20 in female). The five petals are heart shaped (with the point towards the center of the flower) and are deeply cleft into two lobes; the petals form a fringe that makes a distinct collar around the opening to the floral tube. Female flowers have five emergent styles (rarely four or six); male flowers have 10 yellow stamens that barely emerge from the collar at the center of the flower. (Other local Silene sp. have three styles.) The fruit is brown, urn-shaped, and has 10 small teeth on the distal end (or five pairs if you prefer). (Five or six teeth total in S. noctiflora, S. stellata, and S. vulgaris.) Evening campion evolved a dioecious life history (separate male and female plants) about 11 million years ago. (Other species in the genus have "perfect" flowers with both male and female organs present and functional in the flowers.) The evolution and genomic history of the sex-determining Y chromosome (XX = female, XY = male) has been of considerable interest for the light it shines on the evolution of Y chromosomes in both plants and animals (Moraga et al. 2025. Science 387:630–636; Akagi et al. 2025. Science 387:637–643).

Silene

Silene latifolia

evening campion [Blooms: May-Aug]

Royal catchfly's bright red flowers are restricted to the upper regions of the plant as clusters arising from the tips of stems. Individual flowers are 3/4-1" across, have an elongate (1"), narrow, cylindrical calyx with 10 longitudinal ridges, five narrow red or scarlet petals that flare from the tip of the calyx, and stamens with gray anthers that protrude well beyond the opening to the calyx. There are three corkscrew-shaped, threadlike styles that develop after the stamens have matured; like the stamens, they protrude well outside the calyx opening. The fruit is a 1/2" long capsule, open at the top, with 6-8 spreading teeth around the lip. Royal catchfly is hummingbird and butterfly pollinated.

Silene

Silene regia

royal catchfly [Blooms: Jul-Aug]

Starry campion's main stem ends in a panicle of flowers up to 8" long and 3-4" across; additional panicles may arise on the lateral stems on robust plants. Flowers occur in groups of 2-3 on the ends of long, slender branches; the flowers rest directly on short (1/4") pedicels. Individual flowers are 3/4" across. Each flower consists of a bell shaped, light green, fuzzy calyx nearly as broad as it is long with five broad, triangular teeth on its free end; five white petals with a narrow base and a "fringe" of 8-12 narrow lobes on their free ends; 10 stamens with slender white filaments that extend well outside the throat; and three white styles. The styles emerge from a large, green, globular ovary in the middle of the flower. After fertilization, the calyx gets broader (nearly triangular in profile) as the ovary expands into a spherical green ball; the fruit is brown, urn-shaped, and has six teeth on the distal end. The flowers tend to close in bright midday light. They are usually pollinated by moths and, to a lesser extent, bumblebees.

Silene

Silene stellata

starry campion [Blooms: Jun-Jul]

Rosinweed produces a panicle of sunflower-like blooms from the top of the stem and the tops of side stems (if any). Individual flowerheads are 2-3" across; they consist of 12-35 yellow, pistillate (female) ray florets and 70-200+ functionally male disc florets. The green bracts beneath the flowerheads are teardrop-shaped, partially overlapping in two or three layers, and sometimes covered with glandular hairs. The petals of the ray florets have two prominent longitudinal grooves and a tiny apical notch. Only the ray florets can produce seeds — a split style protrudes from a short tube at the base of the ray floret. The central disk is initially green but becomes yellow as the disk florets open; they are yellow, tubular, with five triangular lobes at their free end, with a column of brown stamens with the tips of very long, (non-functional) styles protruding from the tubular corolla. The fruit is a flat seed that develops from the ray florets. (The curious arrangement where only the ray florets can produce seeds while the disk florets are functionally male is true throughout the genus Silphium. The opposite is true in the true sunflowers, genus Helianthus.)

Silphium

Silphium integrifolium

rosinweed [Blooms: Jul-Sep]

Compass plant flower stalks are very hairy, have scattered sessile, triangular bracts, extend vertically multiple feet, and bear 3-4" yellow, sunflower-like composite flowers. There are 20-35 yellow ray florets (e.g., petals) and 100-200+ yellow, tubular disk florets. Behind the flowerheads are two or three layers of very hairy, green, lance-like bracts with sharply pointed, recurved tips. For unknown reason (it just IS), only the ray florets are fertile; the disc florets bear only functional stamens (with a non-functional style protruding through the middle of the stamen). (True throughout the genus Silphium. The opposite is true in the sunflowers, genus Helianthus, where the ray florets are functionally male and the disk florets are fertile, functional females.)

Silphium

Silphium laciniatum

compass plant [Blooms: Jun-Sep]

A cup plant may have 10-30 flowerheads in bloom at one time. Behind the flowerhead are two or three layers of green bracts; the outer layer is egg shaped with a triangular, pointed, recurving tip. Each sunflower-like flowerhead is about 3-4" across; it has 18-40 yellow ray florets (e.g., the petals) and 85-150 yellow disk florets. The ray florets have a bifurcated but functional style protruding from a short yellow tube at the base of the petal; the disc florets are initially green but transform into yellow, tubular corollas with a non-functional style protruding through the fully-functional stamen. (True throughout the genus Silphium. The opposite is true in the sunflowers, genus Helianthus, where the ray florets are functionally male and the disk florets are fertile, functional females.) The cup-like leaves and square stem distinguish cup plants from other prairie wildflowers.

Silphium

Silphium perfoliatum

cup plant [Blooms: Jul-Sep]

Prairie dock's hairless flower stalks produce a panicle of buds and flowerheads. The flowerheads are 2-3" across, with 15-30 bright yellow ray florets with pointed tips and two grooves running the length of the petals; a yellow central disc about 1/4-1/3 the width of the flower contains 120-140 disk florets, green when still buds but bright yellow when open. Behind the flowerhead are two or three layers of green, egg or teardrop shaped, hairless bracts. The ray florets have a bifurcated but functional style protruding from a short yellow tube at the base of the petal; the disc florets have tubular corollas with a non-functional style protruding through the stamen. (True throughout the genus Silphium. The opposite is true in the sunflowers, genus Helianthus, where the ray florets are functionally male, and the disk florets are fertile, functional females.)

Silphium

Silphium terebinthinaceum

prairie dock [Blooms: Jun-Sep]

Wild mustard's upper stems end in racemes of 1/2" wide yellow flowers. Each flower has four yellow petals, six yellow stamens, and a single green pistil. After fertilization, the pistil elongates into a thin, cylindrical silique (seedpod), 1/2-1.5" long, filled with tiny (2 mm diameter) brown or black seeds; the distal third of the silique is a ridged or square "beak". The seeds are reported to remain dormant but viable in the soil for up to 60 years.

Sinapis

Sinapis arvensis

wild mustard [Blooms: May-Jun]

Hedge mustard flowers occur at the tips of long, slender flower stalks that curve upward; the flower stalks typically arise in abundance at the top of the plant. Each flower is small (1/4" across) with four greenish-yellow sepals, four rectangular yellow petals with rounded tips, a single stout style, and six stamens. The fruit is a very slender, cylindrical seedpod (silique) lower on the flower stalk than the blooming flowers; the siliques are 3/8-3/4" long, erect, oriented parallel to and often appressed to the flower stalk, with the remains of the style on their tips. The siliques turn light brown when mature and split open to release the seeds. The short (<3/4"), straight, appressed seedpods and the upward-curving, sparse flower stalks with only a few blooms open at any one time distinguish hedge mustard from other local mustards.

Sisymbrium

Sisymbrium officinale

hedge mustard [Blooms: May-Jul]