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The upper stems of marsh yellow-cress end in one or more 2-8" long racemes of flowers; racemes may also develop from leaf axils. Flowers in bloom are restricted to the tip of the raceme; cylindrical seedpods (siliques) oriented perpendicular to the stem decorate the lower regions of the raceme. Individual flowers are tiny (about 3 mm long) with four greenish-yellow oblong sepals, four yellow spatulate petals, 4-6 stamens with yellow or light brown anthers, and a stout style with a two-lobed stigma. After fertilization, 3-10 mm long, dumbbell-shaped, hairless, green siliques containing two rows of tiny seeds develop. The siliques bear the remnant of the style on one end and 1/2" long pedicels on the other.

Rorippa

Rorippa palustris

marsh yellow-cress [Blooms: Jun-Aug]

Sweet coneflower flowerheads are 2-3.5" across, occurring as single blooms on the tips of stems or branches. A flowerhead is comprised of numerous 1/2" long medium green, lance-shaped phyllaries, downy with short hairs; 10-20 bright yellow, oblong, sterile ray florets with a tiny notch at their tips; and a deep brown hemisphere of 200-400 disk florets (the central disc). The disk florets have a tubular, dark reddish-purple corolla; bifurcated stigmas that have sharp tips; with the purplish-brown anthers adhering side by side around the deep purple styles. Sweet coneflower is distinctive among the Jackson Park coneflowers with its chocolate brown disk and bright yellow, elongate-oval petals with a shallow notch at their tips.

Rudbeckia

Rudbeckia subtomentosa

sweet coneflower [Blooms: Jul-Aug]

Wild mustard's upper stems end in racemes of 1/2" wide yellow flowers. Each flower has four yellow petals, six yellow stamens, and a single green pistil. After fertilization, the pistil elongates into a thin, cylindrical silique (seedpod), 1/2-1.5" long, filled with tiny (2 mm diameter) brown or black seeds; the distal third of the silique is a ridged or square "beak". The seeds are reported to remain dormant but viable in the soil for up to 60 years.

Sinapis

Sinapis arvensis

wild mustard [Blooms: May-Jun]

Hedge mustard flowers occur at the tips of long, slender flower stalks that curve upward; the flower stalks typically arise in abundance at the top of the plant. Each flower is small (1/4" across) with four greenish-yellow sepals, four rectangular yellow petals with rounded tips, a single stout style, and six stamens. The fruit is a very slender, cylindrical seedpod (silique) lower on the flower stalk than the blooming flowers; the siliques are 3/8-3/4" long, erect, oriented parallel to and often appressed to the flower stalk, with the remains of the style on their tips. The siliques turn light brown when mature and split open to release the seeds. The short (<3/4"), straight, appressed seedpods and the upward-curving, sparse flower stalks with only a few blooms open at any one time distinguish hedge mustard from other local mustards.

Sisymbrium

Sisymbrium officinale

hedge mustard [Blooms: May-Jul]

The axils of the middle and upper leaves of woodland goldenrod produce small clusters of 1-12 flowerheads; the stem tip may also develop a small (3" long, 1.5" across) panicle of flowerheads. Individual flowerheads are tiny (3 mm) consisting of 4-5 ray florets with yellow, oblong-elliptical "petals" and 3-6 disc florets with golden yellow, tubular corollas with five tiny lobes. At the base of each flowerhead are three overlapping series of light green, elongate-elliptical bracts (phyllaries). The fruit is a small, bullet-shaped seed 1-2 mm long with a tuft of white hairs 3-5 mm long. In Illinois, only woodland goldenrod and broad-leafed goldenrod (Solidago flexicaulis) produce clusters of flowerheads from the axils of the middle and upper leaves; however, the latter has coarsely serrated leaves with petioles and should be easy to distinguish from woodland goldenrod.

Solidago

Solidago caesia

woodland goldenrod [Blooms: Sep-Oct]

The tip of broad-leaved goldenrod's stem produces a cylindrical inflorescence (a raceme or a narrow panicle) 1.5-5" long and about a third as wide; smaller racemes (1/2-1.5" long, smaller than the leaves) arise from the axils of the upper leaves. Individual flowerheads are about 1/4" across with 3-4 yellow ray florets asymmetrically deployed around the flowerhead and 4-8 yellow disk florets. The base of the flowerhead is surrounded by 3-5 series of unequal, narrowly oblong, light green phyllaries; the outermost phyllaries are 1-2 mm long while the innermost are 3.7-4.7 mm long. Broad-leaved goldenrod prefers high-quality, moist, shaded woodlands. In Illinois, only broad-leaved goldenrod and woodland goldenrod (Solidago caesia) produce clusters of flowerheads from the axils of the middle and upper leaves. Broad-leaved goldenrod has unusually large flowers for a goldenrod and the ray florets are not evenly distributed around the flowerhead. In addition, woodland goldenrod has smooth or finely serrated leaf margins (rather than coarsely-toothed leaves) and no petioles (the leaves are sessile) — it should be easy to distinguish from broad-leaved goldenrod.

Solidago

Solidago flexicaulis

broad-leaved goldenrod [Blooms: Jul-Sep]

The wand-shaped, nodding inflorescence of field goldenrod arises from the tip of the stem; it is 3-7" long and widest in the middle, tapering towards either end; the flowerheads are all on one side of the flower branches. Flowerheads are each 1/4" across consisting of 4-10 yellow ray florets and 3-10 yellow disk florets. The bracts (phyllaries) surrounding the base of the flowerhead are in three overlapping series; they are pale greenish yellow, covered in short, white hairs. The fruit is a dry seed 0.5-2 mm long with an attached tuft of light brown hairs. Field goldenrod can be distinguished from other goldenrods by (1) its diminutive size, (2) the inflorescence that nods to one size, (3) leaflets present in leaf axils (4) short, fine hairs on both leaves and stems, and (5) the grayish cast of the leaves.

Solidago

Solidago nemoralis

field goldenrod [Blooms: Aug-Oct]

Showy goldenrod's inflorescence is a showy, cylindrical panicle up to 12" long arising from the tip of the stem; the flowering stems are erect or curve upwards and the flowerheads occur on all sides of the flowering stems. (Unlike most goldenrods, where the flowering stems curve outward and downward and the flowerheads are on one side of the flowering stems.) Each flowerhead is about 1/4" across consisting of 4-10 large (for a goldenrod) pistillate ray florets (no stamens) and 6-16 perfect disk florets (both stamens and style). Behind the flowerhead are 3-4 series of overlapping bracts (phyllaries), each with a prominent midvein. The fruit is a 2 mm long seed with dark brown longitudinal grooves and a tuft of light brown hairs. Showy goldenrod is distinguished from other goldenrods by the character combination of: (1) stem leaf margins smooth (not toothed), (2) wing-like leaflets in axils of upper leaves, (3) the stems of the inflorescence erect or curve upward (not out and down), and (4) stems and leaves largely or completely hairless except in the inflorescence.

Solidago

Solidago speciosa

showy goldenrod [Blooms: Jul-Oct]

Elmleaf goldenrod's stem (and sometimes upper leaf axils) produces an open, airy inflorescence (a panicle) at its tip that may be a foot tall and wide with arching, widely-spreading branches. The flowerheads rest on very short stems (nearly sessile) along one side of the major branches, facing upward; on the opposite side of the branches are small (1") bracts. Both stems and branches in the inflorescence are covered with hairs. The flowerheads are 1/8-1/4" across with 3-6 yellow, pistillate ray florets (no stamens) and 4-7 yellow, perfect disk florets (both stamens and a style). Surrounding the base of the flowerheads are 16-18 bracts (phyllaries) in 2-3 overlapping series.

Solidago

Solidago ulmifolia

elmleaf goldenrod [Blooms: Jul-Oct]

Purple meadow-rue is a native perennial that is listed as "vulnerable" in Illinois. The species is found in most of the lower 48 states in the U.S. but is notably absent from states on the Atlantic and Pacific coasts (however, it is present along the Gulf of Mexico). Purple meadow-rue is dioecious — some plants produce only staminate (male) flowers, each with around a dozen stamens; other plants bear only pistillate (female) flowers, each with about ten pistils. The male flowers are the showier of the two. Petals are absent (the plant is wind-pollinated). Both the narrow, whitish-grey, thread-like pistils and the yellow-green stamens (with pale yellow anthers) droop from 4-5 light green sepals (that may drop off). Fertilized female flowers develop a single, spindle-shaped fruit that is 1/4" long, elongated, sharply pointed with a remnant style, 4-6 raised ribs, and slightly concave along one side. The fruit is initially green, turning brown with maturity. Early meadow-rue (Thalictrum dioicum) is similar to purple meadow-rue but has light purplish-green stems and leaves with 3-9 (typically five) rounded lobes and long petioles on the upper leaves.

Thalictrum

Thalictrum dasycarpum

purple meadow-rue [Blooms: May-Jun]

Early meadow-rue's main stem terminates in a panicle up to a foot long and wide; each branch in the panicle has a corymb of about five flowers at its tip. Early meadow-rue is dioecious — some plants produce only staminate (male) flowers, each with 10+ stamens; other plants bear only pistillate (female) flowers, each with up to 15 pistils. Both male and female flowers droop from their stems, but the male flowers have a longer droop than do the female flowers. Both male and female flowers are about 1/2" long and 1/4" across with 4-5 pale green sepals with white margins (which often fall off). Neither male nor female flowers have petals; pollination is via the wind. Fertilized female flowers produce ellipsoid, dry seeds that are pointed at both ends; the surface bears strong ribbing. Early meadow-rue prefers shady woodlands. It has more lobes on its leaves (usually 3-5), and the lobes are more rounded than purple meadow-rue (Thalictrum dasycarpum), the other Thalictrum species in Jackson Park, does (2-3 pointed lobes).

Thalictrum

Thalictrum dioicum

early meadow-rue [Blooms: Apr-May]

Field pennycress's central stem and the upper side stems produce racemes of flowers at their tips. The flowers bloom at the tip of the raceme, which continues to elongate to 3-8" long and to produce new buds; the prominent, disk-shaped fruits develop lower on the raceme. Individual flowers are about 1/8-1/4" across on a 1/2" pedicel; there are four green sepals with whitish edges, four white, oval petals, six yellowish-green stamens (three on each side of the flattened ovary), and a single short style. The fruit is a 1/2" wide flattened, winged disc with a distinct U-shaped notch at the apex; the fruits are borne on the flower pedicels, which spread perpendicular to the stem. Each seedpod has two compartments, each compartment containing several dark brown seeds. Other mustards have long, cylindrical seedpods; the nearly-circular seedpod of field pennycress is unique among Jackson Park mustards. Young field pennycress plants are edible but become very bitter at flowering. Cows that feed on field pennycress produce "tainted" milk that tastes like garlic.

Thlaspi

Thlaspi arvense

field pennycress [Blooms: Apr-Jun]

In corn speedwell, single flowers 1/8" wide develop at the tips of inflorescences that arise from the axils of the middle and upper leaves; the flowers are nearly sessile. The inflorescence continually elongates, producing new flowers and a series of bracts along its length. The most basal bracts resemble the stem leaf whose axil hosts the inflorescence; the upper bracts are alternate, 3-10 mm long, and narrowly lance-shaped with long hairs fringing the bract margins. Each flower consists of four hairy, green, lance-like sepals/bracts longer than the corolla lobes; a blue to blue-violet corolla with four petal-like lobes, the upper three lobes slightly larger than the lower forth lobe; two stamens with white anthers; and a pistil with a single style. Dark lines (nectar guides) radiate from the throat of the corolla. Each flower only lasts a single day. The fruit is a 1/8" long and wide, tan, heart-shaped seed capsule, somewhat flattened, with two compartments containing 10-30 yellowish, ovoid seeds. There are 16 species of the genus Veronica reported from Illinois. I have only seen three (Veronica arvensis, Veronica polita, and Veronica serpyllifolia) in Jackson Park but, given their minute size and inconspicuousness, it seems likely that more remain to be found.

Veronica

Veronica arvensis

corn speedwell [Blooms: Apr-Jun]

ird's-eye speedwell (Veronica persica) flowers are solitary, arising from the axils of upper leaves on 1/2-3/4" long, hairy pedicels with a leaf-like bract at their bases. Individual flowers are about 8 mm across, consisting of a calyx with four lance-shaped sepals that are fused at their bases and are shorter than the corolla lobes, and a tubular corolla with four lobes ("petals") that are round to oval. The corolla is bluish toward the edges and whitish centrally, with dark blue lines radiating from the center. One of the four lobes is slightly smaller (and whiter) than the other three, so the flower is bilaterally symmetrical. A pistil with a single 2-3 mm long style arises from the center of the flower; two white stamens with dark blue anthers flank the pistil. Fertilized flowers produce a two-lobed fruit nestled in the remnants of the calyx, each lobe containing 5-11 small seeds; the fruit is a laterally compressed heart shaped capsule that is longer than tall with a distinct shallow V-shaped groove where the remnants of the style can be seen.

Veronica

Veronica persica

bird's-eye speedwell [Blooms: May]

Wayside speedwell's flowers are 1/4" across with a green, hairy calyx with four triangular lobes as long as the petals, projecting between them in the spaces between petals; a short corolla with four spreading lobes, three the same size (one on either side, blue to bluish-white with dark blue lines, and the third dark blue with blue-violet lines) and one about half as wide, pale blue or white with light blue lines. The blue lines function as nectar guides. The base of all four lobes is bright white, but the center of the flower is greenish. There are two white stamens with white anthers (backed by a dark blue "U" shape) and a single white pistil emerging from the throat of the corolla. The fruits are heart-shaped capsules with two chambers covered with glandular hairs, each containing 8-12 seeds. The flowers and leaves of wayside speedwell are similar to those of bird's-eye speedwell (Veronica persica), but the calyx lobes of bird's-eye speedwell do not project beyond the petals. There are 16 species of the genus Veronica reported from Illinois. I have only seen three (Veronica arvensis, Veronica polita, and Veronica serpyllifolia) in Jackson Park but, given their minute size and inconspicuousness, it seems likely that more remain to be found.

Veronica

Veronica polita

wayside speedwell [Blooms: Apr-Jun]