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Early meadow-rue's main stem terminates in a panicle up to a foot long and wide; each branch in the panicle has a corymb of about five flowers at its tip. Early meadow-rue is dioecious — some plants produce only staminate (male) flowers, each with 10+ stamens; other plants bear only pistillate (female) flowers, each with up to 15 pistils. Both male and female flowers droop from their stems, but the male flowers have a longer droop than do the female flowers. Both male and female flowers are about 1/2" long and 1/4" across with 4-5 pale green sepals with white margins (which often fall off). Neither male nor female flowers have petals; pollination is via the wind. Fertilized female flowers produce ellipsoid, dry seeds that are pointed at both ends; the surface bears strong ribbing. Early meadow-rue prefers shady woodlands. It has more lobes on its leaves (usually 3-5), and the lobes are more rounded than purple meadow-rue (Thalictrum dasycarpum), the other Thalictrum species in Jackson Park, does (2-3 pointed lobes).

Thalictrum

Thalictrum dioicum

early meadow-rue [Blooms: Apr-May]

Field pennycress's central stem and the upper side stems produce racemes of flowers at their tips. The flowers bloom at the tip of the raceme, which continues to elongate to 3-8" long and to produce new buds; the prominent, disk-shaped fruits develop lower on the raceme. Individual flowers are about 1/8-1/4" across on a 1/2" pedicel; there are four green sepals with whitish edges, four white, oval petals, six yellowish-green stamens (three on each side of the flattened ovary), and a single short style. The fruit is a 1/2" wide flattened, winged disc with a distinct U-shaped notch at the apex; the fruits are borne on the flower pedicels, which spread perpendicular to the stem. Each seedpod has two compartments, each compartment containing several dark brown seeds. Other mustards have long, cylindrical seedpods; the nearly-circular seedpod of field pennycress is unique among Jackson Park mustards. Young field pennycress plants are edible but become very bitter at flowering. Cows that feed on field pennycress produce "tainted" milk that tastes like garlic.

Thlaspi

Thlaspi arvense

field pennycress [Blooms: Apr-Jun]

Slender stinging nettle plants are dioecious (unisexual) with male (staminate) and female (pistillate) flowers on separate plants. Flowers occur in knobby clusters (panicles) that arise from the leaf axils. Neither pistillate nor staminate flowers have any petals. Staminate flowers have four light green, identical sepals cupping four short stamens. Pistillate flowers also have four sepals but two sepals are narrowly lance-shaped, about a millimeter long, and free, alternating with two sepals that are 1.4-1.8 mm long and cupped around the ovary; styles are absent but the single stigma is long and feathery. The fruit is a flattened, oval to egg-shaped, seed less than 1/16" long. Stinging nettles (aka, tall nettle, slender stinging nettle) have a complicated biogeographic history. According to USDA Plants (https://plants.sc.egov.usda.gov/) and ITIS.gov, the subspecies "Urtica dioica gracilis" is native, while the subspecies "Urtica dioica dioica" is an exotic that is native to Europe and Asia. Thus, you may see "stinging nettle" listed as either "native" or "exotic." Both subspecies are perennials, and both occur in Illinois, although "gracilis" is most common; most features are the same for both subspecies. Wikipedia says these subspecies are actually separate species, an opinion based on a 2014 paper in Phytotaxa (162: 61-83) that analyzed molecular characters; Kew Garden's Plants of the World Online (https://powo.science.kew.org/) and iNaturalist also accept Urtica gracilis as a valid species. I've chosen to use "Urtica gracilis" here if only to avoid further proliferation of long scientific names in this presentation.

Urtica

Urtica gracilis

slender stinging nettle [Blooms: Jul-Aug]

In corn speedwell, single flowers 1/8" wide develop at the tips of inflorescences that arise from the axils of the middle and upper leaves; the flowers are nearly sessile. The inflorescence continually elongates, producing new flowers and a series of bracts along its length. The most basal bracts resemble the stem leaf whose axil hosts the inflorescence; the upper bracts are alternate, 3-10 mm long, and narrowly lance-shaped with long hairs fringing the bract margins. Each flower consists of four hairy, green, lance-like sepals/bracts longer than the corolla lobes; a blue to blue-violet corolla with four petal-like lobes, the upper three lobes slightly larger than the lower forth lobe; two stamens with white anthers; and a pistil with a single style. Dark lines (nectar guides) radiate from the throat of the corolla. Each flower only lasts a single day. The fruit is a 1/8" long and wide, tan, heart-shaped seed capsule, somewhat flattened, with two compartments containing 10-30 yellowish, ovoid seeds. There are 16 species of the genus Veronica reported from Illinois. I have only seen three (Veronica arvensis, Veronica polita, and Veronica serpyllifolia) in Jackson Park but, given their minute size and inconspicuousness, it seems likely that more remain to be found.

Veronica

Veronica arvensis

corn speedwell [Blooms: Apr-Jun]

ird's-eye speedwell (Veronica persica) flowers are solitary, arising from the axils of upper leaves on 1/2-3/4" long, hairy pedicels with a leaf-like bract at their bases. Individual flowers are about 8 mm across, consisting of a calyx with four lance-shaped sepals that are fused at their bases and are shorter than the corolla lobes, and a tubular corolla with four lobes ("petals") that are round to oval. The corolla is bluish toward the edges and whitish centrally, with dark blue lines radiating from the center. One of the four lobes is slightly smaller (and whiter) than the other three, so the flower is bilaterally symmetrical. A pistil with a single 2-3 mm long style arises from the center of the flower; two white stamens with dark blue anthers flank the pistil. Fertilized flowers produce a two-lobed fruit nestled in the remnants of the calyx, each lobe containing 5-11 small seeds; the fruit is a laterally compressed heart shaped capsule that is longer than tall with a distinct shallow V-shaped groove where the remnants of the style can be seen.

Veronica

Veronica persica

bird's-eye speedwell [Blooms: May]

Wayside speedwell's flowers are 1/4" across with a green, hairy calyx with four triangular lobes as long as the petals, projecting between them in the spaces between petals; a short corolla with four spreading lobes, three the same size (one on either side, blue to bluish-white with dark blue lines, and the third dark blue with blue-violet lines) and one about half as wide, pale blue or white with light blue lines. The blue lines function as nectar guides. The base of all four lobes is bright white, but the center of the flower is greenish. There are two white stamens with white anthers (backed by a dark blue "U" shape) and a single white pistil emerging from the throat of the corolla. The fruits are heart-shaped capsules with two chambers covered with glandular hairs, each containing 8-12 seeds. The flowers and leaves of wayside speedwell are similar to those of bird's-eye speedwell (Veronica persica), but the calyx lobes of bird's-eye speedwell do not project beyond the petals. There are 16 species of the genus Veronica reported from Illinois. I have only seen three (Veronica arvensis, Veronica polita, and Veronica serpyllifolia) in Jackson Park but, given their minute size and inconspicuousness, it seems likely that more remain to be found.

Veronica

Veronica polita

wayside speedwell [Blooms: Apr-Jun]

Thyme-leafed speedwell's upper stems host clusters (racemes) of flowers 2-7" long; flowers may either be opposite or alternate along the stem. Each flower is 1/8-1/4" across with a short (1/4"), slender petiole; at the base of the petiole is an oblong, hairless, leafy bract also about 1/4" long with smooth margins. (Thus, the flowers effectively arise from leaf axils.) Individual flowers consist of a green calyx with four triangular lobes shorter than the petals; a short corolla with four oval petal-like, lobes (the lowest one smaller than the other three); two stamens with purple or white anthers; and a green ovary with a single style. The lobes (petals) are white or pale blue, usually with dark blue lines radiating from the flower's center on the upper three lobes. The fruit is a laterally flattened, heart-shaped capsule 1/4" across with two chambers and a fringe of glandular hairs. When mature, the chambers split open, releasing 50-72 tiny seeds. There are 16 species of the genus Veronica reported from Illinois. I have only seen three (Veronica arvensis, Veronica polita, and Veronica serpyllifolia) in Jackson Park but, given their minute size and inconspicuousness, it seems likely that more remain to be found.

Veronica

Veronica serpyllifolia

thyme-leafed speedwell [Blooms: Apr-?]

At the top of the Culver's root plant are floral spikes (racemes) up to 10" long covered with small (1/4" long) white, tubular flowers. Individual flowers consist of a short (3 mm long), green calyx with five deep lobes (the lower two lobes longer than the upper three); a tubular corolla with four lobes (the lower two longer than the upper two); two stamens with yellow or reddish brown anthers; and a single style with a tiny stigma. The stamens and stigma protrude well beyond the lip of the flower. The flowers bloom from the base of the floral spike successively towards the tip. The fruit is a 2.5-4.5 mm long ovoid capsule with four apical teeth. The capsules contain 10-30 half millimeter long, ovoid seeds. Culver's root is very popular with a wide diversity of bees.

Veronicastrum

Veronicastrum virginicum

Culver's root [Blooms: Jun-Aug]