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European spindletree flowers (usually less than 7) occur in clusters (cymes) with long peduncles; they have four sepals, four petals, four stamens with white anthers, and a single stout style. The greenish-white or yellow petals on the flowers appear to be very narrow, half the width of the petals of Hamilton's spindletree. (In reality, the petals are curled into a U-shape towards the stem so they only appear narrow.) The sepals are pale green, about half the length of the petals, and bent back beneath the flower. The fruit is pink, not red; the arils covering the seeds are orange, not red or scarlet. European spindletree blooms later in the spring in Jackson Park than Hamilton's spindletree.

Euonymus

Euonymus europaeus

European spindletree [Blooms: Apr-Jun]

Hamilton's spindletree produces clusters of about seven flowers on ~1.5" peduncles that grow out of the leaf axils. Individual flowers are <1/2" across with four white, elongate-oval petals with rounded tips, four stamens with dark purple (approaching black) anthers, and a large green ovary with a single, stout green style. The fruit is a four chambered, deeply-grooved, rose-colored capsule ~1/2" across containing four scarlet, fleshy arils, each surrounding an ellipsoidal, dark brown seed.

Euonymus

Euonymus hamiltonianus

Hamilton's spindletree [Blooms: Maay-Jun]

Black bindweed flowers develop in singletons or pairs from the leaf axils or directly from the stem. Flowers are 1/8-1/4" long; they have 3-5 greenish-white, white, or light purple petal-like tepals, the outer three tepals with a ridge or keel down the midline on the outer surface. Six to eight stamens with purple anthers surround fused green styles with a dome-shaped stigma in the middle of the flower; the flower usually opens only partially (like a tulip). The fruit is three-sided, with a tubercle-covered surface, about 1/4" long, black when fully mature. Black bindweed is very similar to climbing false buckwheat (Fallopia scandens).

Fallopia

Fallopia convolvulus

black bindweed [Blooms: Jun-Jul]

Forsythia flowers occur singly or in clusters of 2-3 arising from the axils of leaf buds; the flowers have short (1/8-1/4") pedicels. The flowers consist of four green 1/4" long sepals fused at their bases; a bell to saucer-shaped corolla that also has four deep, strap-shaped, brilliant yellow lobes; two stamens that are fused to the base of the corolla tube; and a single style with a pair of ascending, hemispherical terminal lobes (the stigmas). Forsythia is distylous, producing either short (3-5 mm) stamens in combination with long (5-7 mm) pistils or long (6-7 mm) stamens in combination with short (3 mm) pistils. Each of these flower morphs is restricted to a given plant; forsythia is an obligate outcrossing species. Fruits are rarely produced in North American forsythia, possibly because the complementary flower morph is rarely in close proximity to a given plant. However, forsythia can propagate asexually via cuttings.

Forsythia

Forsythia suspensa

forsythia [Blooms: Mar-Apr]

Green ash is dioecius; each tree produces only either male (staminate) or female (pistillate) flowers (which are wind pollinated). The flowers open in mid-spring, before the leaves emerge. Male flowers are produced in compact clusters that envelop short sections of the second-year twigs. Each staminate flower is yellowish green to greenish purple and less than 1/8" (3 mm) across, consisting of a tiny, four-lobed, tubular calyx and a pair of purplish stamens. Petals are absent. Female flowers are also produced on the second-year twigs, in this case in small panicles. Each female flower consists of a tiny four-lobed, tubular calyx and a pistil with a single, long style; again, petals are absent. Fertilized flowers are replaced by panicles of drooping, spatula-shaped, winged seeds 1-2.5" long and 1/4-1/3" wide that have an ellipsoid swelling at one end containing the seed and a single elongated, membranous wing with a rounded or notched tip. When released in the autumn, the greenish to tan fruits spin around the seed, reducing their rate of fall so they can be carried greater distances by the ambient wind. Green ash is tolerant of shading so can grow in mature forests; it prefers moist soils and can tolerate some flooding.

Fraxinus

Fraxinus pennsylvanica

green ash [Blooms: May?]

Gallant-soldier (aka, small-flowered galinsoga) flowerheads are about 1/4" across; they have 4-8 (usually 5) ray florets with a single minute (about 2x2 mm) white petal notched to produce two or three lobes; the ray florets are distributed around the circumstance of the disc. There are 15-50 golden yellow, tubular disc florets with five short triangular lobes at their apex. Both ray and disc florets are fertile; the ray florets are pistillate while the disc florets are perfect (both pistils and stamens). The fruit (a cypsela) consists of a black, conical seed suspended beneath about five off-white, papery, fringed scales that act as a parachute. Gallant-soldier is very similar to galinsoga (Galinsoga quadriradiata); the latter has larger flowerheads and ray floret petals.

Galinsoga

Galinsoga parviflora

gallant-soldier [Blooms: Aug-Sep]

Galinsoga (aka, Peruvian daisy or shaggy soldier) flowerheads are small (1/4" across) with 4-8 (usually five) widely separated 2.5 mm long, white ray florets with three teeth at their tips. The 15-35 disc florets are yellow. Both ray and disc florets are fertile; the ray florets are pistillate while the disc florets are perfect (both pistils and stamens). The fruit (a cypsela) consists of a black, conical seed suspended beneath about five off-white, papery, fringed scales that act as a parachute. A poisonous plant (Tridax procumbens) with nearly identical flowers (but distinctly different leaves) is (luckily) restricted to FL and TX in the U.S. so not likely to show up in Jackson Park. The name Galinsoga ciliata, sometimes still used for Galinsoga quadriradiata, is invalid (ITIS); according to the USDA Plants database, C. ciliata has been superceded by either G. quadriradiata or G. urticifolia.

Galinsoga

Galinsoga quadriradiata

galinsoga [Blooms: Jun-Oct]

Cleavers' flowers are tiny (2-3 mm across), star-shaped, with two green carpels covered with hooked hairs joined at the base of the flower; four white, pointed petals; four stamens; and two styles. The fruit is a spherical capsule formed from the carpels, covered with stiff, hooked hairs, each carpel containing a single seed.

Galium

Galium aparine

cleavers [Blooms: Apr/May-?]

Ground ivy flowers are 1/2" long and arise from the leaf axils; they are purplish, trumpet shaped, with a notched upper and lower (the largest) lobe and two small side lobes; the "throat" of the lower lobe is hairy. There are four stamens (two long, two short) with purple to blue anthers, and a single pistil with a bifurcated purple style. The corolla of the flowers is cupped by a hairy, green, tubular calyx that has 15 ridges distributed around the calyx running longitudinally, and five triangular teeth at the apex of the calyx. The fruit is a nutlet that nestles in the calyx.

Glechoma

Glechoma hederacea

ground ivy [Blooms: Apr-May]

Dame's rocket flowers occur in clusters (racemes) of up to 30. Each flower is 3/4-1" across and long, with a long, narrow, tubular, pinkish-green calyx composed of four long narrow sepals. The calyx opens through a narrow throat into four rounded, pinkish-purple (sometimes white) petals; the six pale yellow stamens and the multiple styles poke to various degrees out of the throat. Stems, leaves, pedicles, and the flower calyxes are all downy or hairy. The fruits are long (up to 4"), narrow, ascending, downy cylinders (siliques, typical of mustards) filled with flattened, oblong seeds. The flowers are very phlox-like, but phlox flowers have five petals, and dame's rocket only has four; phlox leaves are opposite on the stem (not alternate) and are narrower.

Hesperis

Hesperis matronalis

dame's rocket [Blooms: Apr-May]

Wild hydrangea canes produce a flat terminal panicle 3-6" across with 100-500 flowers of two kinds — small, fertile flowers filling most of the area and a few, sterile pseudo-flowers around the margin that are 3-4 times larger than the flowers in the center and seem to function only as an advertisement for pollinators. The central, fertile flowers are about 1/4" across with a green calyx made of five separate sepals; five greenish-white or cream petals about 1/8" long; 8-10 stamens with long filaments and oblong, white anthers; and a pistil bearing two styles with knob- or club-like stigmas. The sterile, outer pseudo-flowers have 3-4 fused petal-like, white bracts (or sepals; sources differ) with a central structures that visually is reminiscent of stamens or styles; the pseudo-flowers persist long after the inner flowers have been fertilized and have produced a two compartment seed pod that is ribbed and bears a pair of tiny horns (the styles) on its upper end. Wild hydrangea is relatively rare in northern Illinois; in Jackson Park it prefers wooded area by walking paths.

Hydrangea

Hydrangea arborescens

wild hydrangea [Blooms: Jun-Aug]

Panicled hydrangea (Hydrangea paniculata) is an exotic perennial native to China and Japan that may grow as either a shrub or a tree. Neither Natureserve Explorer nor the USDA Plants database acknowledge the presence of panicled hydrangea in Illinois, but the Morton Arboretum and the University of Illinois Extension both explicitly do. Panicled hydrangea is widely sold in nurseries; commercial cultivars reach 10-15 feet high and wide. It definitely occurs in Jackson Park, near our native species — wild hydrangea (Hydrangea arborescens) — but I have not seen it in direct competition with wild hydrangea. The inflorescence is a conical panicle typically covered with two types of flowers — large, nominally-sterile flowers and smaller, bisexual flowers — scattered across the surface of the panicle. The "sterile" flowers are white or greenish white with typically 4-5 (3 in Jackson Park) elliptical to round lobes (actually sepals) 1/2-1" long and 1/3-2/3" across (8-15 mm). In Jackson Park, at least half the nominally sterile flowers bear a small, budlike structure in their center that exhibits three lobes (petals?), have 3-6 distinct stamens with small anthers, and a two-lobed central structure that is probably a reduced pistil. It is not clear whether either the pistils or stamens are functional. The bisexual flowers usually have five tiny (less than 1 mm long and wide) sepals, five petals 3-4 mm long and 0.1-0.3 mm wide (which often fall off the flower), 2-5 mm long stamens, and an ovary with 2-4 spreading styles (two in Jackson Park specimens) that are 2-3.5 mm long and are fused at their bases. After fertilization, the petals and stamens wither and fall off; the remainder of the flower forms an oblong, 3-5 mm long, seed capsule. The easiest ways to tell panicled hydrangea (H. paniculata) from wild hydrangea (H. arborescens) is: (1) the inflorescence is conical or pyramid-shaped in panicled hydrangea but a flat disc or shallow dome in wild hydrangea, and (2) the showy, "sterile" flowers in panicled hydrangea are scattered evenly across the surface of the inflorescence in panicled hydrangea, but are restricted to the margins of the inflorescence of wild hydrangea (rather like the inflorescence of the much more common but unrelated American cranberry, Viburnum opulus var. americanum). Wild hydrangea is much more common than panicled hydrangea in Jackson Park.

Hydrangea

Hydrangea paniculata

panicled hydrangea [Blooms: Jul]

Black walnut has separate male and female flowers on the same tree (often the same branch). The male florets occur on 3-6" long, dangling, green, cylindrical catkins that arise from the leaf axils of one-year-old branchlets; the florets are 1/8" across and are comprised of a short, tubular calyx with 2-6 lobes and 15-40 stamens with greenish-yellow or purplish anthers. The female florets occur in groups of 2-5 on short spikes that arise from the tips of the current year's new branchlets; these florets are also about 1/8" across but are comprised of a short, tubular calyx with four lobes, a stout, green ovary covered with sticky hairs, and a pair of stout, bushy styles with red-tinged stigmas. The female florets are wind pollinated; the trees are self-fertile if cross-pollination does not occur. Fertilized female florets develop into globoid nuts about 2.5" long and 2" across that initially retain the styles; the nuts are solitary or in pairs. The thick husk turns from green to yellowish-brown to black as the fruit matures. The nuts cannot germinate unless they are exposed to 90-120 days of cold winter temperatures.

Juglans

Juglans nigra

black walnut [Blooms: May?-Jun]

American water-willow produces long (up to 6") peduncles from some of the upper leaf axils that have clusters of flowers on their tips. Individual flowers are 3/4" across with a short, tubular, 1/4" long, green calyx with five teeth and a tubular corolla with four lobes — an upper lobe with a shallow V-notch, two widely spreading lateral lobes, and one lower lobe at right angles to the other three. Both the lateral and lower lobes are folded downward along their midlines, producing a U-shaped cross-section. All of the lobes are white or pale purple with a darker splotch on the end of the upper lobe and dark purple spots along the midline of the lower lobe and at the base of the lateral lobes. There are two stamens, each with a dark brown pair of anthers posed in front of the upper corolla lobe and a single, narrow, white style running along the midline off the upper lobe and protruding beyond its upper boundary.

Justicia

Justicia americana

American water-willow [Blooms: Jun-Aug]

Wood nettle plants may be either monoecious (separate male and female flowers on the same plants) or unisexual (only male or only female flowers). Male flowers are produced from branching cymes 4" long that arise from the leaf axils; female flower are produced from branching cymes, also 4" long, localized near the apex of the stem. Male flowers are small (1/8" across), consisting of five narrow, greenish-white sepals and five stamens; there are no petals but there is a vestigial pistil. Female flowers are about the same size (1/8" across) with four green sepals (two larger, two smaller) that look like tiny leaves with undulating edges and an ovary with a long, white, vertically-oriented J-shaped style that tapers to a point. The female flowers are wind-pollinated. The fruit is circular in outline but very flattened, green turning black with maturation, on a short winged stalk; the remains of the style are attached at the free end of the fruit.

Laportea

Laportea canadensis

wood nettle [Blooms: Aug-Sep]

Field peppergrass tems produce racemes of flowers near their apex; flowers near the top of the raceme are the youngest with older flowers (and later seed pods) further down the inflorescence. The flowers have a calyx of green or purple-green, lance-like sepals, four white, spoon-shaped petals, six stamens, and a single style. The fruits are flattened pods oval in outline with the remnants of the style protruding from a notch in the end (rather like field pennycress, Thlaspi arvense); each seedpod contains two seeds. This is an easy plant to overlook if only because it look like the weed that it is.

Lepidium

Lepidium campestre

field peppergrass [Blooms: May-?]

Peppergrass (aka, Virginia pepperweed) is a weedy, 1.5' tall, rather bushy plant. The flower spikes (racemes) are 2-4" long with tiny (<1/8") white apical flowers forming a larger inflorescence. The flowers have four oblong green or tan sepals with a white edge, half the length of the petals, alternating with four white, triangular petals. There are two stamens with yellow anthers and a stubby (0.2 mm tall) style on a green ovary. Below the flowers, the flower spike is covered with 1/8" wide seedpods on (relatively) long stalks, reminiscent of a bottlebrush; the seedpods are oval with an apical notch (much like half-sized versions of field pennycress (Thlaspi arvense) seedpods) containing two seeds. This species is native to the U.S. The stubby cylindrical inflorescences are and the triangular petals alternating with shorter sepals are diagnostic and distinguish pepper grass from field peppergrass (Lepidium campestre).

Lepidium

Lepidium virginicum

peppergrass [Blooms: Oct-Nov]

American bugleweed flowers form in whorls of the axils of the middle and upper leaves. Individual flowers are 1/8" across, white with pinkish-purple spots, with a calyx about as long as floral tube with five teeth; the distal end of the short tubular corolla has five sharply-pointed, narrowly-triangular lobes. The two stamens with purple anthers and single style project well outside the mouth of the corolla. The fruits are four, flat-topped nutlets that form in the cup of the calyx. The deep, narrow lobes of the lower leaves are a characteristic of this species; they are not present in the other Lycopus species in Jackson Park.

Lycopus

Lycopus americanus

American bugleweed [Blooms: Jul-Sep]