Panicled hydrangea (Hydrangea paniculata) flowers. Top right: the inflorescence of panicled hydrangea. Note that the large, sterile (?) flowers occur across the entire domed surface of the panicle. Bottom right: the underside of an inflorescence showing the branching structure of the panicle. Top left: three of the large, sterile (?) flowers of panicled hydrangea. Note the large sepals, the remnants of (infertile) stamens, and the bulbous structure in the center of the flowers (petals?). Bottom left: three tiny fertile flowers, probably recently fertilized. The five sepals and two styles are easily visible; the petals and stamens have both probably been shed.
Panicled hydrangea produces large, pyramidal clusters of flowers, quite distinct from the typical flattened or slightly domed panicles of wild hydrangea (although not of some commercial cultivars of that species). The inflorescence is a conical panicle typically covered with two types of flowers — large, nominally-sterile flowers and smaller, bisexual flowers — scattered across the surface of the panicle. The “sterile” flowers are white or greenish white with typically 4-5 (3 in Jackson Park) elliptical to round lobes (actually sepals) 1/2-1″ long and 1/3-2/3″ across (8-15 mm). In Jackson Park, at least half the nominally sterile flowers bear a small, budlike structure in their center that exhibits three lobes (petals?), have 3-6 distinct stamens with small anthers, and a two-lobed central structure that is probably a reduced pistil. It is not clear whether either the pistils or stamens are functional. The bisexual flowers usually have five tiny (less than 1 mm long and wide) sepals, five petals 3-4 mm long and 0.1-0.3 mm wide (which often fall off the flower), 2-5 mm long stamens, and an ovary with 2-4 spreading styles (two in Jackson Park specimens) that are 2-3.5 mm long and are fused at their bases. After fertilization, the petals and stamens wither and fall off; the remainder of the flower forms an oblong, 3-5 mm long, seed capsule.
Panicled hydrangea (Hydrangea paniculata) leaves. The upper surface (top image) and underside (lower image) of the same leaf of a panicled hydrangea.
Panicled hydrangea (Hydrangea paniculata) is an exotic perennial native to China and Japan that may grow as either a shrub or a tree. Neither Natureserve Explorer nor the USDA Plants database acknowledge the presence of panicled hydrangea in Illinois, but the Morton Arboretum and the University of Illinois Extension both explicitly do. Panicled hydrangea is widely sold in nurseries; commercial cultivars reach 10-15 feet high and wide. It definitely occurs in Jackson Park, near our native species — wild hydrangea (Hydrangea arborescens) — but I have not seen it in direct competition with wild hydrangea. The easiest ways to tell panicled hydrangea (H. paniculata) from wild hydrangea (H. arborescens) is: (1) the inflorescence is conical or pyramid-shaped in panicled hydrangea but a flat disc or shallow dome in wild hydrangea, and (2) the showy, “sterile” flowers in panicled hydrangea are scattered evenly across the surface of the inflorescence in panicled hydrangea, but are restricted to the margins of the inflorescence of wild hydrangea (rather like the inflorescence of the much more common but unrelated American cranberry, Viburnum opulus var. americanum). Wild hydrangea is much more common than panicled hydrangea in Jackson Park.

